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Acytolepis Toxopeus, 1927

Acytolepis Toxopeus, 1927; Tijdschr. Ent. 70 (3/4) : 271, 288, TS: Polyommatus puspa Horsfield
Acytolepsis[sic?] ; Hemming, 1967, Bull. Br. Mus. Nat. Hist. (Ent.), Suppl.: 25
Acytolepis (Polyommatini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 449
Acytolepsi (Polyommatini) ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

600x400 (~104Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~101Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~79Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar
600x400 (~49Kb) India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Acytolepis puspa (Horsfield, [1828])

Peninsular India, Himalayas, Assam, Ceylon, Burma, Philippines, Nicobars, New Guinea. See [About maps] Common Hedge Blue
Polyommatus puspa Horsfield, [1828]; Descr. Cat. lep. Ins. Mus. East India Coy (1): 67, TL: Java
Cyaniris splendens Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29) : 444, TL: Larut Hills, Perak
Cyaniris puspa sania Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 285
Cyaniris puspa contilia Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 285, TL: Sumbawa, Sumba
Cyaniris puspa calata Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 286, TL: Flores
tymbria (Fruhstorfer, 1922) (Lycaenopsis); in Seitz, Gross-Schmett. Erde 9 : 870
umenonis (Matsumura, 1931) (Lycaenopsis); Insecta Matsumurana 6: 44
haani (Kalis, 1938) (Lycaenopsis); Ent. Rdsch. 55: 198
Lycaenopsis puspa ; [BIR, 274]
Celastrina puspa ; [BOW: pl. 175, f. 20, 21]
Acytolepis puspa ; [MRS, 679]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

600x400 (~59Kb) underside India: Subansiri, 31.10.2005, Photo © Kedar Tokekar

600x400 (~75Kb) underside India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

Larva on Glochidion fortunei, Rhododenron spp., Distylium racemosum, Rosa multiflora, Rosa wichuraina, R. centifolia, Prunus zippeliana, Glochidion obovatum, Myrica rubra, Quercus phillyraeoides, Celtis sinensis, Astilbe thunbergii [MRS], Schleichera oleosa, Hiptage benghalensis, Xylia dolabriformis [BMP], Viburnum, Combretaceae, Cycadaceae, Shorea, Enkianthus, Lyonia, Rhododendron, Vaccinium, Antidesma, Breynia, Bridelia, Glochidion, Calliandra, Cylista, Paracalyx, Pellophorum, Xylia, Quercus, Distylum, Hiptage, Myrica, Aruncus, Chaenomeles, Malus, Photinia, Prunus, Rosa, Spiraea, Dimocarpus, Euphoria, Litchi, Sapindus, Schleichera, Astilbe, Celtis Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

A. p. puspa

A. p. cagaya (C. & R. Felder, [1865])

Philippines (Luzon) Lycaena cagaya C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 278, pl. 34, f. 11-13
Cyaniris puspa ottonis Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 287, TL: Palawan

A. p. lambi (Distant, 1882)

Polyommatus lambi Distant, 1882; Ann. Mag. nat. Hist. (5) 10 (57) : 245, TL: Malay Peninsula
Cyaniris imperatrix Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29) : 444, TL: Thailand
Acytolepis puspa lambi, Butterflies in Indo-China [Yutaka Inayoshi]
Acytolepis puspa lambi ; [BMP: 240, pl. 35, f. 44-46]

A. p. puspinus (Kheil, 1884)

Nias puspa puspinus (Kheil, 1884) (Plebeius); Die Rhopalocera der Insel Nias: 30, f. 37-39, TL: Nias

A. p. kuehni (Röber, 1886)

Sulawesi Plebeius kühni Röber, 1886; Corr.-Bl. Ent. Ver. Iris 1 (3) : 60, pl. 4, f. 29 (partim), TL: E.Sulawesi
Plebeius kuehni Röber, 1886
Lycaenopsis puspa martini Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 870, TL: Sulawesi

= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

Lycaenopsis puspa hyllus Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 870, TL: Banggai

= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

Cyanirus[sic] kuehni kalawari Ribbe, 1926; Ent. Mitt. 15 : 89, TL: Sulawesi, Kalawara

= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

Cyaniris kuehni bangkaiensis Ribbe, 1926; Ent. Mitt. 15 : 89, TL: Banggai

= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

Lycaenopsis puspa kuehni ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2) : 438
Cyaniris kuhni[sic] ; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2) : 287
Acytolepis puspa kuehni ; Toxopeus, 1928, Tijdschr. Ent. 71 : 231; Eliot & Kawazoé, 1983, Butts. Lycaenopsis group: 180; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
Acytolepis puspa kühni ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54, f. 23-24

A. p. cyanescens (de Nicéville, 1890)

Nicobar Is. Cyaniris cyanescens de Nicéville, 1890; in Marshall & de Nicéville, Butts India Burmah Ceylon 3 : 103

A. p. prominens (de Nicéville, 1890)

Cyaniris cyanescens prominens de Nicéville, 1890; in Marshall & de Nicéville, Butts India Burmah Ceylon 3 : 103

A. p. dammae (Heron, 1894)

Timor, Damar, Wetar Cyaniris dammae Heron, 1894; Ann. Mag. nat. Hist. (6) 14 (80) : 104, TL: Damma I.
Acytolepis puspa dammae ; [BAUR, 383]

A. p. timorensis (Butler, 1900)

Timor Cyaniris timorensis Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29) : 443, TL: Timor
Acytolepis puspa timorensis ;

A. p. myla (Fruhstorfer, 1909)

Formosa Cyaniris puspa myla Fruhstorfer, 1909; Ent. Zs. 23 (25) : 116, TL: Formosa
myla (Matsumura, 1929) (Cyaniris); Insecta Matsumurana 3 (2/3) : 104 (preocc. Fruhstorfer)
formosana (Bethune-Baker, 1914) (Celastrina); Ent. Mitt. 3 (4) : 127 [?]
Acytolepis puspa myla ; [MRS, 679]

Larva on Glochidion acuminatum, G. lanceolatum, G. rubrum, Bridelia balansae, Euphoria longan, Sapindus mukorossi, Vaccinium oldhamii, V. samllii, Enkyathe perulatus, Lyonia ovalifolia, Viburnum dilatatum, Prunus padus, Chaenomeles speciosa, Malus domestica, Spiraea cantoniensis, Aruncus dioicus, Astilbe thunbergii [MRS]

A. p. hermagoras (Fruhstorfer, 1910)

Hainan Cyaniris puspa hermagoras Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 284, TL: Hainan
Acytolepis puspa hermagora ; [MRS, 679]

A. p. gisca (Fruhstorfer, 1910)

NE.Afghanistan - Burma, Thailand, Cambodia, Laos, Vietnam, S.China, Yunnan Cyaniris puspa gisca Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 285, TL: Sikkim
Lycaenopsis puspa gisca f. artena Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 25, TL: India
Lycaenopsis puspa telis Fruhstorfer, 1917; Archiv Naturg. (A) 82 (1) : 25, TL: Andaman
Celastrina puspa barneyi Corbet, 1941; Hongkong Nat. 10 (3/4): 149, TL: Hong Kong
Acytolepis puspa gisca, Butterflies in Indo-China [Yutaka Inayoshi]
Acytolepis puspa gisca ; [MRS, 679]; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)

A. p. hermione (Fruhstorfer, 1905)

Bawean puspa hermione (Fruhstorfer, 1905) (Cyaniris); Soc. Ent. 20 (1) (1): 2
Cyaniris puspa hermione ; Fruhstorfer, 1905, Ent. Zs. 19 (12): 64, f. 10

A. p. bazilana (Fruhstorfer, 1910)

Philippines (Mindoro, ...?) Cyaniris puspa bazilana Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 286, TL: Bazilan
Cyaniris puspa georgi Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 286, TL: Camiguin de Mindanao
sabis (Fruhstorfer, 1917) (Lycaenopsis); Archiv Naturg. (A) 82 (1) : 25

A. p. darmis (Fruhstorfer, 1917)

Kai, Aru? Acytolepis puspa darmis ; [BAUR, 383 (text)]

A. p. mygdonia (Fruhstorfer, 1917)

Sumatra, Borneo puspa mygdonia (Fruhstorfer, 1917) (Lycaenopsis); Archiv Naturg. (A) 82 (1) : 26

A. p. roberi (Fruhstorfer, 1917)

Goram puspa roberi (Fruhstorfer, 1917) (Lycaenopsis); Archiv Naturg. (A) 82 (1) : 27, TL: Goram
Acytolepis puspa roberi ; [BAUR, 383 (text)]

A. p. volumina (Fruhstorfer, 1922)

Lycaenopsis volumina Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 869, Lfg. (302)

A. p. deronda (Fruhstorfer, 1922)

Sula Is. Lycaenopsis puspa deronda Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 871, TL: Sula Is
Lycaenopsis puspa var. Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2) : 438
Acytolepis puspa deronda ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162

A. p. vandeldeni Toxopeus, 1929

Sumatra puspa vandeldeni Toxopeus, 1929; Tijdschr. Ent. 72 (3/4) : 207

A. p. felderi Toxopeus, 1927

Ceylon Acytolepis puspa felderi Toxopeus, 1927; Tijdschr. Ent. 70 (3/4) : 294

A. p. ishigakiana (Matsumura, 1929)

Cyaniris ishigakiana Matsumura, 1929; Insecta Matsumurana 3 (2/3) : 104, TL: Okinawa

A. p. siporana (Riley, [1945])

Mentawi Is. puspa siporana (Riley, [1945]) (Celastrina); Trans. R. ent. Soc. Lond. 94 (2): 256

Acytolepis lilacea (Hampson, 1889)

Nilgiris, Palnis, Coorg. See [About maps] Hampson's Hedge Blue
Cyaniris lilacea Hampson, 1889; J. asiat. Soc. Bengal 57: 356
?Cyaniris puspa lilacina[sic] ; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2) : 285 (missp.)
Lycaenopsis lilacea ; [BIR, 274]
Lycaenopsis puspa lilacea ;

A. l. lilacea

A. l. moorei (Toxopeus, 1926)

Ceylon Lycaenopsis moorei Toxopeus, 1926; Treubia 8: 365

A. l. indochinensis Eliot & Kawazoé, 1983

Burma, Thailan, Laos Acytolepis lilacea indochinensis Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 187, f. 135, 268, 284, TL: Vang Vien, Laos
Acytolepis lilacea indochinensis, Butterflies in Indo-China [Yutaka Inayoshi]

Acytolepis ripte (Druce, 1895)

Borneo. See [About maps] Cyaniris ripte Druce, 1895; Proc. zool. Soc. Lond. 1895 (3) : 574, pl. 32, f. 11

Acytolepis najara (Fruhstorfer, 1910)

Sulawesi. See [About maps] Cyaniris najara Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2) : 287 (partim, ♂), TL: Sulawesi, Bua-Krang, 5000ft
Acytolepis najara ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162

21.4.2012 (5)

References:

[About maps]

Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

[BAUR]

D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)

[BIR]

Wynter-Blyth, M. A., 1957; Butterflies of the Indian Region; (1982 Reprint)

[BMP]

Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2

[BOW]

Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X

[Kedar Tokekar]

India

[LAMAS-2005]

Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)

[MRS]

Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2

[NHM]

Natural History Museum; The Global Lepidoptera Names Index

[Yutaka Inayoshi]

A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM

Some related literature:

Bethune-Baker, 1914

H. Sauter's Formosa-Ausbeute : Ruralidae (Lep.) Ent. Mitt. 3 (4) : 123-127

Butler, 1900

A List of the Species of Cyaniris, a well-known Group of the Family Lycaenidae Ann. Mag. nat. Hist. (7) 5 (29) : 441-451

Cassidy, 1995

On the Lycaenopsis Group of Genera (Lepidoptera, Lycaenidae) of the Sulawesi Region Trans. lepid. Soc. Japan 46 (2): 45-62

Chapman, 1909

A review of the species of the lepidopterous genus Lycaenopsis Feld. (Cyaniris auct. nec Dalm.) on examination of the male ancilliary appendages Proc. zool. Soc. Lond. 1909 (2) : 419-476

Distant, 1882

Undescribed Rhopalocera from the Malay Peninsula Ann. Mag. nat. Hist. (5) 10 (57) : 245-248

Druce, 1895

A Monograph of the Bornean Lycaenidae Proc. zool. Soc. Lond. 1895 (3) : 556-627, pl. 31-34

Eliot, 1956

New and little known Rhopalocera from the Oriental Region Bull. Raffles Mus. 27: 32-38, 3 figs

Eliot, 1973

The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 373-505, 162fgs, pl. 1-6

Eliot & Kawazoé, 1983

Blue butterflies of the Lycaenopsis group. British Museum (Natural History) Butts. Lycaenopsis group: 309 pp., 560 figs

C. & R. Felder, [1865]

Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Vers.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)

Fruhstorfer, 1905

Neue Lepidopteren von Bawean und Nachbarinseln Soc. Ent. 20 (1) (1): 2-4 (1 April), (2): 9-10 (15 April)

Fruhstorfer, 1905

Neue Rhopaloceren aus dem Indo-Australischen Gebiet Ent. Zs. 19 (9): 45-47, f. 1-5 (1 June), (10): 51 (8 June), (11): 55-57, f. 6-9 (15 June), (12): 63-64, f. 10 (22 June)

Fruhstorfer, 1909

Lepidopterologisches Pêle-Mêle. [XVI]. Neues aus Formosa Ent. Zs. 23 (25) : 116-117 (18 September)

Fruhstorfer, 1910

Neue Cyaniris-Rassen und Übersicht der bekannten Arten Stettin ent. Ztg. 71 (2) : 282-305 (15 March)

Fruhstorfer, 1917

Revision der Lycaenidengattung Lycaenopsis auf Grund morphologischer Vergleiche der Klammerorgane Archiv Naturg. (A) 82 (1) : 1-42, 2 pls. (January)

Fruhstorfer, 1922

Gattungen (8-19): Parelodina - Niphanda in Seitz, Gross-Schmett. Erde 9 : 856 (14 February 1921), : 857-864 (5 January 1922), : 865-872 (21 January, 1922), : 873-880 (15 April 1922), : 881-888 (18 April 1922), : 889-896 (16 May 1922), : 897-901 (16 March 1923)

Hemming, 1967

Generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera) Bull. Br. Mus. Nat. Hist. (Ent.), Suppl.: -

Heron, 1894

A Visit to Damma Island, East India Archipelago. Lepidoptera Ann. Mag. nat. Hist. (6) 14 (80) : 101-107

Horsfield, [1828]

Descriptive Catalogue of the Lepidopterous Insects contained in the Museum of the Horourable East-India Company, illustrated by coloured figures of new species Descr. Cat. lep. Ins. Mus. East India Coy (1): 1-80, pl. 1-4 (1828), (2): 81-144, pl. 5-8, (expl.) [1-2] (1829)

Huang, 2003

A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55: 3-114

de Nicéville in Marshall & de Nicéville, 1890

The Butterflies of India, Burmah and Ceylon, a descriptive Handbook of all the known Species of rhopalocerous Lepidoptera inhabiting that Region, with Notices of allied Species occurring in the neighbouring Countries along the Border, with numerous Illustrations. Vol. 3 Butts India Burmah Ceylon 3 : 1-503, pl. 25-29

Matsumura, 1929

New butterflies from Japan, Korea and Formosa Insecta Matsumurana 3 (2/3) : 87-107, pl. 4

Pagenstecher, 1897

in Kuekenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und in Borneo. Lepidopteren Abh. Senckenb. naturf. Ges. 23: 353-467, 3pls

Ribbe, 1926

Neue Lycaenenformen hauptsächlich von Celebes Ent. Mitt. 15 : 78-91

Riley, [1945]

Spolia Mentawiensia: Rhopalocera, Lycaenidae and Riodinidae Trans. R. ent. Soc. Lond. 94 (2): 247-271

Röber, 1886

Neue Tagschmetterlinge der Indo-Australischen Fauna Corr.-Bl. Ent. Ver. Iris 1 (3) : 45-72, pl. 2-5

Toxopeus, 1927

Eine Revision der javanischen, zu Lycaenopsis Felder und verwandten Genera gehörigen Arten. (Mit Einführung von 6 neuen Genera, Beschreibung von 2 neuen Species un 5 neuen Subspecies). Lycaenidae Australasiae II Tijdschr. Ent. 70 (3/4) : 232-302, pl. 1, 27 text figs.

Toxopeus, 1928

Eine Revision der javanischen, zu Lycaenopsis Felder und verwandten Genera gehörigen Arten. Lycaenidae Australasiae II Tijdschr. Ent. 71 : 179-265, pl. 1, 29 text figs.

Toxopeus, 1929

Beschreibung einiger Schmetterlinge (Riodinidae und Lycaenidae) von Pulu Weh bei Sumatra. (Lycaenidae Australasiae V) Tijdschr. Ent. 72 (3/4) : 204-214

Vane-Wright & de Jong, 2003

The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.