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Melitaea Fabricius, 1807
Melitaea Fabricius, 1807; Magazin f. Insektenk. (Illiger) 6: 284, TS: Papilio cinxia Linnaeus
Schoenis Hübner, [1819]; Verz. bek. Schmett. (2): 28, TS: Papilio delia Denis & Schiffermüller
Cinclidia Hübner, [1819]; Verz. bek. Schmett. (2): 29, TS: Papilio phoebe Denis & Schiffermüller
Melinaea Sodoffsky, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6) : 80 (unn. repl. Melitaea Fabricius, 1807, preocc. Melinaea Hübner, 1816), TS: Papilio cinxia Linnaeus
Didymaeformia Verity, 1950; Le Farfalle diurn. d'Italia 4: 89, 90, TS: Papilio didyma Esper
Melitaea, NSG Voucher Specimen [Wahlberg]
Melitaea (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165
Didymaeformia (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166
Cinclidia (Melitaeini) ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166
Melitaea (Nymphalidae) ; [BRU2, 59]
Melitaea (Melitaeina) ; [Wahlberg]; [AfrL]

The "didyma" species-group

Melitaea didyma (Esper, 1778)
NW.China. See [About maps] Spotted Fritillary Red-Band Fritillary Roter Scheckenfalter Doncella timida Przeplatka didyma
Papilio didyma Esper, 1778; Die Schmett. Th. I, Bd. 1 (7): pl. 41, f. 3
?Papilio didyma Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): pl. 61, f. 1, TL: Bavaria
Papilio cytheris Mueschen, 1781
Papilio athulia Fabricius, 1787
orientalis Herrich-Schäffer, 1856; (preocc.)
alpina Staudinger, 1861; (nom. nud.)
occidentalis Staudinger, 1861; (nom. nud.)
meridionalis Staudinger, 1861; (nom. nud.)
dalmatina Staudinger, 1861; (nom. nud.)
graeca Staudinger, 1861; (nom. nud.)
rossica Staudinger, 1861; (nom. nud.)
meridionalis Staudinger, 1870
occidentalis Staudinger, 1781
dalmatina Staudinger, 1870
Melitaea didyma f. problematica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 357, pl. 27, f. 11-12
Melitaea didyma, NSG Voucher Specimen [Wahlberg]
Melitaea didyma, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea didyma, Moths and Butterflies of Europe [MBE]
Melitaea didyma, Europaische Schmetterlinge [Mario Maier]
Melitaea didyma, Captain's European Butterfly Guide [Simon Coombes]
Melitaea didyma ; [H&R, 116]
Didymaeformia didyma ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea didyma ; [BOW: pl. 3, f. 2-3]; [EBW]
Melitaea (didyma) didyma ; [BRU2: 59, pl. 38, f. 1-12, 25-26]
Melitaea didyma ; [MRS, 586]; [Otakar Kudrna]
[didyma-1fu.jpg] 500x316 (~61Kb) upperside female Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes [didyma-1mu.jpg] 500x325 (~44Kb) upperside male Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes [didyma-1v.jpg] 450x432 (~38Kb) underside Axat, Pyrenees Orientales 24/07/96, Photo © Simon Coombes
[didyma-2l.jpg] 597x400 (~49Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [didyma-2p.jpg] 371x550 (~36Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch [didyma-2u.jpg] 597x400 (~52Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Plantago, Veronica, Viola, Scrophularia, Linaria, Dianthus, (etc.) [BRU2, 59]


SEU, CEU, W.Siberia, Caucasus, Transcaucasia kasanskyi Krulikovsky, 1901
marginata Ksienschopolsky, 1912; (preocc.)
niesiolowskii Bryk, 1940
uralicola Bryk, 1940
crimitaea Bryk, 1940
maledescripta Bryk, 1940
pontifex Bryk, 1940
itkolensis Alberti, 1969
Melitaea didyma didyma ; [BRU2, 59]
M. d. kirgisica Bryk, 1940
Tarbagatai, Saur, Tian-Shan ?geminella Bryk, 1940
Melitaea didyma kirgisica ; [BRU2, 59]
M. d. turkestanica Sheljuzhko, 1929
Alai Melitaea didyma turkestanica Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 354 (repl. Melitaea turanica Staudinger, 1886)
Melitaea turanica Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 231 (preocc.)
Melitaea turkestanica ab. pseudoala Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 355, TL: Kauntshi, Tashkent
Melitaea didyma turkestanica ; [BRU2, 59]
M. d. elavar Fruhstorfer, 1917
Ghissar Melitaea didyma elavar ; [BRU2, 59]
M. d. occidentalis Staudinger, 1861
Spain, N.Africa Melitaea didyma occidentalis Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8, TL: Albarracin, Spain
Melitaea didyma f. mauretanica Oberthür, 1909; Étud. Lépid. Comp. 3 : 243, TL: Algeria, Spain
Melitaea didyma var. meridionalis Staudinger, 1870; Horae Soc. ent. ross. 7 (1870) : 60, TL: Mt. Parnassus, Greece
Melitaea didyma occidentalis f. dalmatina Staudinger, 1870; Horae Soc. ent. ross. 7 (1870) : 60, TL: Greece
Melitaea didyma nisseni Rothschild, 1913; Novit. Zool. 20 (1) : 115, TL: Aļn Draham (Tunisia)
Melitaea didyma interposita Rothschild, 1913; Novit. Zool. 20 (1) : 115, TL: Batna (Algeria)
Melitaea didyma mauretanica f. occasus Verity, 1919; Ent. Rec. J. Var. 31 : 180, TL: [N] Africa
Melitaea didyma proteaoccidentis Verity, 1929; Ent. Rec. J. Var. 41 : 117, TL: Fum Kheney [Foum Kheneg], Mekney [Meknes] (Morocco)
Melitaea didyma africa-felix Bryk, 1940; Folia zool. hydrobiol. 10: 326, pl. 1, f. 5, TL: El-Bayadh, Sebdou (Algeria)
Melitaea didyma nisseni f. gen. tunisa Eisner, 1942; Zool. Meded. Leiden 24 (4): 118, TL: Tunis (Tunisia)
Melitaea didyma meridionalis ; [H&R, 116]
Melitaea didyma occidentalis ; [H&R, 117]; [BMAT: 53, pl. 17, f. 22-36]

Larva on Linaria spp., Scrophularia sp., Antirrhinum sp. [BMAT], (other regions) Mispate orontium, Plantago sp., Plantago amplexicaulis, Melampyrum sp., Verbascum sp., Linaria sagittata [BMAT]


interposita
[interposita-1v.jpg] 600x435 (~41Kb) underside Morocco, Tizi-n-Test, Haut Atlas, Photo © Michel Tarrier


mauretanica
[mauretanica-1uf.jpg] 600x435 (~43Kb) upperside female Morocco, Djebel Lakraa, Rif, Photo © Michel Tarrier [mauretanica-2um.jpg] 600x435 (~28Kb) upperside male Morocco, Itzer, Moyen Atlas, Photo © Jean Delacre

M. d. neera Fischer de Waldheim, 1840
Volga, S.Ural, SW.Siberia Melitaea uralca Bryk, 1940
[neera-1l.jpg] 820x480 (~92Kb) larva A high steppefied right bank of the Shipunikha Rivulet 3 km NE of the village Evsino, Iskitim district, Novosibirsk Province, West Siberia, Russia. 8th June 1996 (on a leaf of Phlomis tuberosa L.), Photo © Oleg Kosterin

Larva on Phlomis tuberosa [Oleg Kosterin]


?M. d. ambra Higgings, 1941
dschungarica Oberthür, 1909; (preocc.)
perplexa Higgins, 1941
manoni Hanus, 1996
ambra ; [BRU2, 59 (?)]

Melitaea interrupta Kolenati, 1846
-. See [About maps] Melitaea interrupta Kolenati, 1846; Meletamata Entomologica 5 : 86, TL: "Caucaso et Transcaucasia"
Melitaea transcaucasica Turati, 1919; Naturalista sicil. 23 (7-12) : 225
= ; [Wahlberg]
caucasi Verity, 1929; (preocc.)
venosata Niesiolowsky, 1937
distinctissima Bryk, 1940
musatscherii Alberti, 1969
Melitaea transcaucasica, NSG Voucher Specimen [Wahlberg]
Didymaeformia transcaucasica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea (didyma) interrupta ; [BRU2: 60, pl. 38, f. 13-18]
NE.Turkey?, Caucasus, Transcaucasus Melitaea interrupta interrupta ; [BRU2, 60]
M. i. kendevana Gross & Ebert, 1975
Kopet-Dagh maculata Gross & Ebert, 1975; (preocc.)
elbursicola Gross & Ebert, 1975
safawides Gross & Ebert, 1975
?wernickei Gross & Ebert, 1975
grossi Koēak, 1980
Melitaea interrupta kendevana ; [BRU2, 60]

Melitaea latonigena Eversmann, 1847
-. See [About maps] Melitaea latonigena Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3) : 66, pl. 1, f. 1-2, TL: [Irkutsk Region, Russia]
atrata Higgins, 1935
mongoligena Bryk, 1940
Melitaea latonigena, Lepidoptera collection [SZMN]
Melitaea latonigena, NSG Voucher Specimen [Wahlberg]
Melitaea didyma latonigena ;
Melitaea didyma latonigera[sic] ; [MRS, 586]
Melitaea (didyma) latonigena ; [BRU2: 60, pl. 37, f. 31-36]
Tuva - Baikal Melitaea latonigena latonigena ; [BRU2, 60]
M. l. altaica Grum-Grshimailo, 1893
Altai latonigena altaica Grum-Grshimailo, 1893; , TL: SW.Altai, Lake Markakol
Melitaea latonigena altaica ; [Oleg Kosterin]; [BRU2, 60]
[altaica-1mfv.jpg] 461x700 (~58Kb) underside male female A subalpine short forb meadow on an opening in an open larch forest on the southern principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets , 2100 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16th July 1998, Photo © Oleg Kosterin [altaica-1mv.jpg] 400x533 (~49Kb) underside male An alpine meadow, at 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 14th July, 1988, Photo © Oleg Kosterin [altaica-2v.jpg] 429x668 (~28Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin
[altaica-2vx.jpg] 768x700 (~76Kb) underside A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998 (on Veronica porphyriana), Photo © Oleg Kosterin [altaica-3fu.jpg] 572x407 (~28Kb) upperside female A larch parkland between the Chikty and Akbul Rivulets, 2000 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 15th July 1998, Photo © Oleg Kosterin [altaica-3mvA.jpg] 503x432 (~21Kb) (aberrative) underside male A steppefied southern slope of small mountains on the Dzhazator River right bank in front of the Akbul Rivulet mouth, between the Yuzhno-Chuiskii mountain range and the Ukok Plateau, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 19th July 1998, Photo © Oleg Kosterin

M. l. polaris Grum-Grshimailo, 1899
Yakutia Melitaea latonigena polaris ; [BRU2, 60]

Melitaea mixta Evans, 1912
-. See [About maps] Melitaea mixta Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 583, TL: "Chit.[ral]"
Didymaeformia mixta ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea didyma mixta ;
Melitaea (didyma) mixta ; [BRU2: 61, pl. 41, f. 1-3]
NW.India Melitaea mixta mixta ; [BRU2, 61]
M. m. nadezhdae Sheljuzhko, 1912
E.Pamirs Melitaea didyma nadezhdae Sheljuzhko, 1912; Dt. ent. Z. Iris 26 (3) : 137, TL: Pamirskij-Post
Melitaea mixta nadezhdae ; [BRU2, 61]
?M. m. problematica Sheljuzhko, ?1912
W.Pamirs Melitaea mixta problematica ; [BRU2, 61]

Melitaea persea Kollar, [1850]
Iran, Asia Minor - Afghanistan, W.Tien-Shan. See [About maps] Melitaea persea Kollar, [1850]; Denkschr. Akad. Wiss. Wien. 1 : 52, TL: "Farsistan" [Shiraz, Iran]
kaschtschenkoi Christoph, 1889
= ; [BRU2, 61]
Melitaea didyma magnacasta Verity, 1929
= ; [BRU2, 61]
Melitaea didyma araratica Verity, 1929
= ; [BRU2, 61]
Melitaea didyma tauricus Belter, 1934
= ; [BRU2, 61]
microtauricus Belter, 1934
= ; [BRU2, 61]
hafiz Higgins, 1941
= ; [BRU2, 61]
afghana Heydemann, 1954
= ; [BRU2, 61]
darius Gross & Ebert, 1975
= ; [BRU2, 61]
pfeifferi Gross & Ebert, 1975
= ; [BRU2, 61]
?wernickei Gross & Ebert, 1975
= ; [BRU2, 61]
Melitaea persea, NSG Voucher Specimen [Wahlberg]
Didymaeformia afghana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Didymaeformia persea ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea persea ; Weiss & Major, 2000, Atalanta 31 (1/2): 123 (note), pl. VII, f. 9-16; [AfrL]; Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note)
Melitaea (didyma) persea ; [BRU2: 61, pl. 44, f. 1-9]
M. p. caucasica Staudinger, 1870
persea caucasica Staudinger, 1870; Horae Soc. ent. ross. 7 (1870) : 59
= Melitaea persea Kollar, [1850]; [BRU2, 61]
caucasica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8 (nom. nud.)
M. p. dogsoni Grose-Smith, 1887
Afghanistan Melitaea dogsoni Grose-Smith, 1887; Ann. Mag. nat. Hist. (5) 20 : 130, TL: Gwashki
= Melitaea persea Kollar, [1850]; [BRU2, 61]
M. p. paphlagonia Fruhstorfer, 1917
persea paphlagonia Fruhstorfer, 1917; Archiv Naturg. (A) 82 (2) : 10
= Melitaea persea Kollar, [1850]; [BRU2, 61]
M. p. montium Belter, 1934
Syria, Lebanon Melitaea montium Belter, 1934; Arb. Morph. Tax. Ent. Berl. 1: 107
= Melitaea persea Kollar, [1850]; [BRU2, 61]
M. p. sargon Hemming, 1932
Iraq persea sargon Hemming, 1932; Trans. ent. Soc. Lond. 80 : 272
= Melitaea persea Kollar, [1850]; [BRU2, 61]
Melitaea persea sargon ; [AfrL]

Melitaea jitka Weiss & Major, 2000
SW.Iran. See [About maps] Melitaea jitka Weiss & Major, 2000; Atalanta 31 (1/2): 123, pl. V, f. 1-4, pl. VI, pl. VII, f. 1-8, TL: SW.Iran,Buyer Ahmed-Kuhgilueye, Yasuj, Dinar Mts., 2600-2700m

Melitaea casta (Kollar, [1848])
Iran. See [About maps] Melitatea casta Kollar, [1848]; Denkschr. Akad. Wiss. Wien. 1 : 52
Melitaea casta, NSG Voucher Specimen [Wahlberg]
Didymaeformia casta ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea casta ; Weiss & Major, 2000, Atalanta 31 (1/2): 126 (note), pl. V, f. 5-8

Melitaea saxatilis Christoff, 1876[1873?]
Iran. See [About maps] Melitaea saxatilis Christoff, 1876[1873?]; Hor. Soc. Ent. Ross. 10: 28
Melitaea fergana var. hyrcana Staudinger, 1882; in Staudinger & Bang-Haas, Berl. ent. Z. 26 (1) : 168
Melitaea saxatilis, NSG Voucher Specimen [Wahlberg]
Didymaeformia saxatilis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea saxatilis ; [BOW: pl. 196, f. 7]

Melitaea deserticola Oberthür, 1909
N.Africa, Tripolitania, Egypt, Lebanon, Transjordan. See [About maps] Desert Fritillary
Melitaea ab. deserticola Oberthür, 1876; Étud. d'Ent. 1 : 25, pl. 3, f. 1, TL: Biskra (Algeria)
Melitaea deserticola, NSG Voucher Specimen [Wahlberg]
Melitaea deserticola, Moths and Butterflies of Europe [MBE]
Melitaea deserticola ; [H&R, 117]
Didymaeformia deserticola ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea deserticola ; [AfrL]
[deserticola-1u.jpg] 600x435 (~35Kb) upperside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre [deserticola-2uf.jpg] 600x435 (~68Kb) upperside female Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre [deserticola-3v.jpg] 600x435 (~59Kb) underside Morocco, Plateau de l'Arid, Haut Atlas, Photo © Jean Delacre
[deserticola-4u.jpg] 600x435 (~53Kb) upperside Morocco, Aļt-Abdallah, Anti-Atlas , Photo © Michel Tarrier

Tunisia, Algeria, Morocco Melitaea didyma harterti Rothschild, 1913; Novit. Zool. 20 (1) : 115, TL: El Hadadra, El Golea, Ghardaia (Algeria)
Melitaea didyma harterti ab. pseudoacraeina Rothschild, 1917; Novit. Zool. 24 (1): 102, TL: El Hadadra (Algeria)
Melitaea didyma harterti ab. nigrofasciata Rothschild, 1917; Novit. Zool. 24 (1): 102, TL: El Hadadra (Algeria)
Melitaea deserticola deserticola ; [BMAT: 54, pl. 17, f. 37-38, pl. 18, f. 1-7]

Larva on Linaria aegyptiaca, Plantago media, Anarrhinum fruticosum, (in Lebanon) Anarrhinum spp. [BMAT]

M. d. macromaculata Belter, 1934
Melitaea deserticola macromaculata ; [AfrL]
M. d. scotti Higgins, 1941
Melitaea abyssinica scotti Higgins, 1941
Melitaea deserticola scotti ; [AfrL]

Melitaea abyssinica Oberthür, 1909
Ethiopia. See [About maps] Didymaeformia abyssinica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea abyssinica ; [BAFR, 213]; [AfrL]

Melitaea didymina Staudinger, 1895
Mongolia. See [About maps] Melitaea didymina Staudinger, 1895; Dt. Ent. Z. Iris 8 (2) : 347
Didymaeformia didymina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)

Melitaea didymoides Eversmann, 1847
-. See [About maps] Melitaea didymoides Eversmann, 1847; Bull. Soc. Imp. Nat. Moscou 20 (3) : 67, pl. 1, f. 3-4, TL: [Kyakhta, Buryatia]
sibirica Staudinger, 1861; (nom. nud.)
sibirica [Heyne], [1893]; (preocc.)
mandschurica Seitz, [1909]; (preocc.)
seitzi Matsumura, 1929
mandschukoana Bryk, 1940
Melitaea didymoides, NSG Voucher Specimen [Wahlberg]
Didymaeformia didymoides ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea didymoides ; [MRS, 586]; [BRU2: 62, pl. 39, f. 19-24]
[didymoides-1mv.jpg] 592x710 (~76Kb) underside male A steppefied meadow at Lake Butyvken, surrounded by a magnificent pine forest within a steppe zone called Tsasucheiskii Bor, 10 km SW of the village Nizhnii Tsasuchei, Onon District, SE Chita Province, Transbaikalia, Siberia, Russia. 4th July 1996, Photo © Oleg Kosterin

S.Tuva?, Transbaikalia, Amur Melitaea didymoides didymoides ; [BRU2, 62]
M. d. yugakuana Matsumura, 1927
S.Ussuri Melitaea didyma yugakuana Matsumura, 1927; Insecta Matsumurana 1 (4) : 164, TL: S.Manchuria, Prov. Yugakujo
Melitaea didymoides yugakuana ; [BRU2, 62]
M. d. pekinensis Seitz, [1909]
N.China Melitaea didymoides pekinensis ; [MRS, 586]; [BRU2, 63]
M. d. latonia Grum-Grshimailo, 1891
C.China Melitaea didymoides latonia ; [MRS, 586]; [BRU2, 62]
M. d. eupatides Fruhstorfer, 1917
Gansu cansicola Bryk, 1940
Melitaea didymoides eupatides ; [BRU2, 63]
M. d. hummeli Bryk, 1940
S.Mongolia Melitaea didymoides hummeli ; [BRU2, 63]

Melitaea ala Staudinger, 1881
Tian-Shan, Dzhungarsky Ala-Tau Mts., S.Altai. See [About maps] Melitaea didyma var. ala Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 288, TL: Lepsa, Ala-Tau
Melitaea ala, NSG Voucher Specimen [Wahlberg]
Didymaeformia ala ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea ala ; [BRU2: 63, pl. 39, f. 10-15]

Larva on Ligularia [BRU2, 63]

S.Altai, Saur Mts., Dzhungarsky Ala-Tau, Burchansarytau Mts., Tyshkantau Mts., Kaiandy-Tau, Horgos R., Alty-Emel Mts. Melitaea ala ab. immodulata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 366, TL: Dzharkent
Melitaea ala allah Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 346, pl. 3, f. 23, TL: "Siberia"
Melitaea ala ; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7) : 171, f. 5-6; Wagner, 1913, Ent. Mitt. 2 (3): 91-93
Melitaea ala ala ; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19 : 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 346
Melitaea ala ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 242-244, pl. 2, f. 4, 10; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: 184-185, pl. 36, f. 5-6
Melitaea ala ala ; Kolesnichenko, 1999, Atalanta 30 (1/4): 91, pl. 6, f. 1-4; [BRU2, 63]
M. a. bicolor Seitz, 1907
Kungey Ala-Tau, Kirgizsky Mts., Suusamyr Valley, Suusamyr Mts., Talassky Ala-Tau, Terskey Ala-Tau, Baidullu Mts., Naryn-Too Mts. Melitaea ala ab. bicolor Seitz, 1907; Gross-Schmett. Erde 1 : 219, pl. 66, f. 7-8, TL: Karagatai Mts.
Melitaea ala ab. latemarginata Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 366, pl. 28, f. 7-8, TL: Naryn
Melitaea ala bicolor ; Sushkin, 1913, Z. wiss. Insektenbiologie 9 (6/7) : 172; Sheljuzhko, 1929, Mitt. Münch. Ent. Ges. 19 : 368; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 347-348; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 244, pl. 6, f. 6, 12; Kolesnichenko, 1999, Atalanta 30 (1/4): 93, pl. 6, f. 5-7; [BRU2, 63]
M. a. zaisana Lukhtanov, 1999
S.Altai Melitaea ala zaisana Lukhtanov, 1999; Atalanta 30 (1/4): 143, TL: Kazakstan, Ust-Kamenogorsk, S.Altai, between Kurtschum and Kalguty, 600m
Misidentified/misapplied:
Melitaea ala; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: pl. 35, f. 5-6
M. a. irtyschica Lukhtanov, 1999
W.Altai Melitaea ala irtyschica Lukhtanov, 1999; Atalanta 30 (1/4): 144, TL: Kazakstan, Ust-Kamenogorsk, Ulbinski, Oktjabrsk, 420m
E.Tian-Shan Melitaea ala sheljuzhkoi ; [BRU2, 63 (?)]
E.Tian-Shan Melitaea ala determinata ; [BRU2, 63 (?)]
?M. a. strandi Bryk, 1940
E.Tian-Shan Melitaea ala strandi ; [BRU2, 63 (?)]

Melitaea kotshubeji Sheljuzhko, 1929
Peter the Great Mts., N.Hazreti-Sho Mts., W.Zaalaisky Mts., S slopes (Alaisky Mts.), SW slopes (Fergansky Mts.), SE slopes (Cathkalsky Mts.) Melitaea ala kotshubeji Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 364, pl. 27, f. 1-4, TL: Touptshek
Melitaea kotshubeji ; [BRU2: 64, pl. 38, f. 31-32, pl. 39, f. 1-3]

Larva on Ligularia thomsonii [BRU2, 65]

Peter the Great Mts., Hazreti-Sho, Zaalaisky Mts. Melitaea didyma var. ala Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 430, TL: Touptshek
Melitaea ala kotshubeji ; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 348
Melitaea chitralensis mod. kotshubeji ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 4, 10
Melitaea ala kotshubeji ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
Melitaea kotshubeji kotshubeji ; Kolesnichenko, 1999, Atalanta 30 (1/4): 96, pl. 6, f. 8-11
Misidentified/misapplied:
Melitaea ala bicolor Seitz; Shchetkin, 1981, Ent. Obozr. Tajikistana: 163-164

Larva on Ligularia thompsonii Shchetkin, 1981, Ent. Obozr. Tajikistana

M. k. bundeli Kolesnichenko, 1999
W.Zaalaisky Mts., S.Alaisky Mts. Melitaea kotshubeji bundeli Kolesnichenko, 1999; Atalanta 30 (1/4): 98, pl. 6, f. 12-13, TL: Alaisky Mts., near Daraut-Kurgan, 3000
Melitaea kotshubeji bundeli ; [BRU2, 65]
M. k. kugarti Kolesnichenko, 1999
northern slopes (E.Alaisky Mts.), Fergansky, Chatkalsky Mts. Melitaea kotshubeji kugarti Kolesnichenko, 1999; Atalanta 30 (1/4): 100, pl. 6, f. 14-17, TL: Fergansky Mts., Kugart river, 2000

Melitaea ninae Sheljuzhko, 1935
Chatkalsky Mts., Pskempsky Mts., Kuraminsky Mts., Talassky Ala-Tau Mts., Vissokoe, Chandalakshsky Mts. Melitaea ala ninae Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25 : 28, TL: Tian-Shan, Tshimgan
Melitaea ala ninae f. magnifica Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25 : pl. 3, f. 5
Melitaea ala ninae f. oblongomaculata Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25 : pl. 3, f. 6
Melitaea ala ninae f. elongatoconfluens Sheljuzhko, 1935; Mitt. Münch. ent. Ges. 25:
Melitaea rosea Higgins, 1938; Entomologist 71: 34, TL: Alie-Ata
Melitaea ala ella-claudia Bryk, 1940; Folia Zoologica et Hydrobiologica 10 (2): 351, pl. 5, f. 49-50, pl. 6, f. 54, TL: Aulie-Ata, Talas Alatau
Melitaea ninae, NSG Voucher Specimen [Wahlberg]
elongotoconfluens ; [BRU2, 64 (missp.?)]
Melitaea ninae ; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: 85, pl. 36, f. 7-8; Kolesnichenko, 1999, Atalanta 30 (1/4): 101, pl. 7, f. 18-21; [BRU2: 64, pl. 39, f. 7-9]
Misidentified/misapplied:
Melitaea pseudoala Sheljuzhko; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 245, pl. 4, f. 12, pl. 16, f. 4
Melitaea pseudoala Sheljuzhko; Higgins, 1955, Trans. R. ent. Soc. Lond. 106 (1): 116
Melitaea pseudoala Sheljuzhko; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166
Melitaea pseudoala Sheljuzhko; Korshunov, 1972, Ent. Obozr. 51 (2): 355
Melitaea pseudoala Sheljuzhko; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48

Melitaea chitralensis Moore, 1901
Kashmir, Chitral. See [About maps] Melitaea chitralensis Moore, 1901; Lepidoptera Indica, 5 : 9, pl. 380, f. 1, TL: Chitral
Melitaea didyma chitralipluvia Verity, 1929; Ent. Rec. J. Var. 41 : 43, TL: Chitral
Melitaea chitralensis, NSG Voucher Specimen [Wahlberg]
Melitaea chitralensis ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 247, pl. 6, f. 3, 9; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); Kolesnichenko, 1999, Atalanta 30 (1/4): 103, pl. 7, f. 22-25

Melitaea enarea Fruhstorfer, 1917
Melitaea didyma enarea Fruhstorfer, 1917; Archiv Naturg. (A) 82 (2) : 11, TL: Garm, Peter the Great Mts.
Melitaea enarea, NSG Voucher Specimen [Wahlberg]
Melitaea enarea ; [BRU2: 64, pl. 38, f. 19-24, 27-30]
Garm, Kamara-su river, Peter the Great Mts., Hazreti-Sho, Darvazky Mts., Shugnansky Mts. Melitaea didyma shugnana Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 355, pl. 27, f. 7-10, TL: Chorag

Melitaea didyma enarea ; Bryk, 1940, Folia Zoologica et Hydrobiologica 10 (2): 333, pl. 1, f. 7, pl. 5, f. 52
Melitaea chitralensis enarea ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7): 248, pl. 6, f. 2; Shchetkin, 1981, Ent. Obozr. Tajikistana: 164-165
Melitaea didyma enarea ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 47
Melitaea chitralensis shugnana ; Tshikolovets, 1997, The Butterflies of Pamir: 118, pl. 28, f. 7-8, pl. 29, f. 7-8, pl. 43, f. 4
Melitaea enarea enarea ; Kolesnichenko, 1999, Atalanta 30 (1/4): 106, pl. 7, f. 26-29
Melitaea enarea shugnana ; [BRU2, 64]
Melitaea enarea enarea ; [BRU2, 64]
M. e. ishkashima Sheljuzhko, 1929
Ishkashimsky Mts. Melitaea didyma ishkashima Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 359, pl. 27, f. 13-14, TL: Kishlak Njut
= Melitaea enarea shugnana Sheljuzhko, 1929; [BRU2, 64]
Melitaea didyma ishkashima ; Tshikolovets, 1997, The Butterflies of Pamir: pl. 28, f. 13, pl. 29, f. 13, pl. 38, f. 8, pl. 39, f. 8
Melitaea enarea ishkashima ; Kolesnichenko, 1999, Atalanta 30 (1/4): 108, pl. 7, f. 30
M. e. permuta Higgins, 1941
Nuratau Mts., Kugitang Mts., Gissarsky Mts., Zeravshansky Mts., Turkenstansky Mts., Karateginsky Mts. Melitaea enarea mod. permuta Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 250, pl. 6, f. 7-8, TL: Samarkand, Hazreth-Sultan
Melitaea permuta, NSG Voucher Specimen [Wahlberg]
Melitaea ala permuta ; Tuzov, 1993, The synonymic list of Butterflies from the ex-USSR: 48
Melitaea enarea permuta ; Kolesnichenko, 1999, Atalanta 30 (1/4): 109, pl. 7, f. 31
Melitaea permuta ; [BRU2: 64, pl. 39, f. 4-6]
M. e. gromenkoi Kolesnichenko, 1999
Fergansky Mts. Melitaea enarea gromenkoi Kolesnichenko, 1999; Atalanta 30 (1/4): 110, pl. 7, f. 32-34, TL: Kirghizia, 15km NW from Dzhalal-Abad, Kara-Kianyr Mts., 1200m
Melitaea enarea gromenkoi ; [BRU2, 64]

Melitaea pseudoala Sheljuzhko, 1929
Turkestan. See [About maps] Melitaea pseudoala Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19 : 355
= Melitaea didyma turkestanica ; [Wahlberg]
Didymaeformia pseudoala ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)

Melitaea sutschana Staudinger, 1892
-. See [About maps] Melitaea sutschana Staudinger, 1892; in Romanoff, Mém. Lépid. 6 : 183, TL: Sutschan-Gebiete
sutschanala Bryk, 1940
transbaicalica Bryk, 1940
Melitaea sutschana, NSG Voucher Specimen [Wahlberg]
Didymaeformia sutschana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea sutschana ; [BRU2: 63, pl. 39, f. 16-18]
[sutschana-1fv.jpg] 410x400 (~37Kb) underside female A steppefied meadow on a southern slope of the Polovinnaya Pad' valley few kilometres of the Argun' River left bank 13 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 28th July 1997 (aberrantly large basal black spots), Photo © Oleg Kosterin [sutschana-2fu.jpg] 410x547 (~53Kb) upperside female A steppefied meadow on the Argun' River left bank terrace 4 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997, Photo © Oleg Kosterin [sutschana-3mv.jpg] 533x400 (~35Kb) underside male A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz.), Photo © Oleg Kosterin
[sutschana-4mu.jpg] 565x490 (~34Kb) upperside male A steppefied southern left slope of the Budyumkan River valley 5 km of its confluence with the Argun River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 26th July 1997 (on Leontopodium conglobatum (Turcz.) Hand.-Mazz), Photo © Oleg Kosterin

Transbaikalia - Amur, Ussuri, Korea, NE.China Melitaea sutschana sutschana ; [BRU2, 63]
M. s. graeseri Gorbunov, 1995
Sakhalin Melitaea sutschana graeseri ; [BRU2, 63]

Melitaea acraeina (Staudinger, 1886)
Fergana Valley Melitaena acraeina Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 233, TL: Kokand and Fergana, Fergana Valley, Uzbekistan
Melitaea acraeina, NSG Voucher Specimen [Wahlberg]
Didymaeformia acraeina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea acraeina ; [BOW: pl. 195, f. 4]; [BRU2: 65, pl. 44, f. 13-15]

Larva on Dodartia orientalis [BRU2, 65]


Melitaea lunulata Staudinger, 1901
-. See [About maps] Melitaea lunulata Staudinger, 1901; in Staudinger & Rebel, Cat. Lepid. palaearct. Faunengeb., 1: 30, TL: Issyk-Kul
Melitaea lunulata, NSG Voucher Specimen [Wahlberg]
Didymaeformia lunulata ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea lunulata ; [BRU2: 65, pl. 43, f. 16-18]
Tian-Shan Melitaea lunulata lunulata ; [BRU2, 66]
M. l. merke Lukhtanov, 1999
W.Tian-Shan Melitaea lunulata merke Lukhtanov, 1999; Atalanta 30 (1/4): 147, TL: Kazakstan, Merke Fl., 2900
Melitaea lunulata merke ; [BRU2, 66]
Misidentified/misapplied:
Melitaea fergana lunulata; Lukhtanov & Lukhtanov, 1994, Herbipoliana, 3: pl. 33, f. 5

Melitaea fergana Staudinger, 1882
Turkestan. See [About maps] Melitaea fergana Staudinger, 1882; in Staudinger & Bang-Haas, Berl. ent. Z. 26 (1) : 168, TL: Alai-Gebirge [Fergana, Turkestan]
Melitaea fergana, NSG Voucher Specimen [Wahlberg]
Didymaeformia fergana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea fergana ; [BRU2: 66, pl. 42, f. 31-42, pl. 43, f. 1-12]
Alai, Transalai Melitaea fergana fergana ; [BRU2, 66]
M. f. maracandica Staudinger, 1882
Ghissarsky Mts. Melitaea fergana var. maracandica Staudinger, 1882; in Staudinger & Bang-Haas, Berl. ent. Z. 26 (1) : 168, TL: Alai (Fergana, Turkestan)
Didymaeformia macarandica[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name) [spell.?]
Melitaea fergana maracandica ; [BRU2, 66]
M. f. darvasika Higgins, 1941
Darvaz Melitaea fergana darvasika ; [BRU2, 66]
M. f. jacobsoni Higgins, 1941
W.Pamirs Melitaea fergana jacobsoni ; [BRU2, 66]
M. f. paradoxa Churkin & Tuzov, 2000
N.Tian-Shan Melitaea fergana paradoxa Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana paradoxa ; [BRU2, 66]
M. f. ketmeana Lukhtanov, 1999
S.Kazakstan, NE.Kirghizia Melitaea fergana ketmeana Lukhtanov, 1999; Atalanta 30 (1/4): 145, TL: Kazakstan, Alma-Ata, Ketmen, Kolshat
Melitaea fergana ketmeana ; [BRU2, 66]
M. f. terskeana Lukhtanov, 1999
Kirghizia Melitaea fergana terskeana Lukhtanov, 1999; Atalanta 30 (1/4): 146, TL: Kirghizia, Terskey-Alatoo, Barskoun-Pass, 3400m
Melitaea fergana terskeana ; [BRU2, 66 (?)]
M. f. oxuana Lukhtanov, 1999
E.Pamir Melitaea fergana oxuana Lukhtanov, 1999; Atalanta 30 (1/4): 147, TL: Tadjikistan, E.Pamir, Sarykol, Dunkeidyk, 3750
M. f. grumi Churkin & Tuzov, 2000
E.Pamir Melitaea fergana grumi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana grumi ; [BRU2, 66]
M. f. vladislavi Churkin & Tuzov, 2000
Inner Tian-Shan (At-Bashi Mts.) Melitaea fergana vladislavi Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana vladislavi ; [BRU2, 66]
M. f. khantengri Churkin & Tuzov, 2000
Inner Tian-Shan (Kaindy Mts.) Melitaea fergana khantengri Churkin & Tuzov, 2000; Helios 1: (33-60) [?]
Melitaea fergana khantengri ; [BRU2, 66]
M. f. splendida Churkin & Tuzov, 2000
Turkestansky Mts. (Pass Kumbel) Melitaea fergana splendida Churkin & Tuzov, 2000; Helios 1: (33-60) [?]

Melitaea infernalis Grum-Grshimailo, 1891
Tian-Shan, Dzhungarsky Alatau. See [About maps] Melitaea infernalis Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 455
Melitaea infernalis, NSG Voucher Specimen [Wahlberg]
Didymaeformia infernalis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea infernalis ; [BRU2: 67, pl. 43, f. 13-15]

Melitaea ambrisia Higgins, 1935
Alai, Ghissar. See [About maps] Melitaea ambrisia Higgins, 1935; Entomologist 68: 112, TL: Alai Mts., Kirgizia
Melitaea ambrisia, NSG Voucher Specimen [Wahlberg]
Didymaeformia ambrisia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea ambrisia ; Churkin, 2000, Atalanta 31 (1/2): 114, pl. IV, f. 1-3; [BRU2: 67, pl. 43, f. 25-30]
Alai Melitaea ambrisia ambrisia ; [BRU2, 67]
M. a. avicaena Churkin, 2000
Turkestansky Mts. (Kumbel Pass) Melitaea ambrisia avicaena Churkin, 2000; Atalanta 31 (1/2): 115, pl. IV, f. 4-6, TL: Uzbekistan, Turkestansky Range, Kumbel Pass, Zaaminsky rs., 3000m
Melitaea ambrisia avicaena ; [BRU2, 67]

Melitaea alraschid Higgins, 1941
-. See [About maps] Melitaea ambrisia alraschid Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 259-260, TL: Persia [= error] [?W Gissar, near Sarmarkand]
Melitaea alraschid ; Churkin, 2000, Atalanta 31 (1/2): 119, pl. IV, f. 7-9; [BRU2: 67, pl. 43, f. 19-24]
Ghissar Melitaea alraschid alraschid ; [BRU2, 67]
M. a. garun Churkin, 2000
Matcha Melitaea alraschid garun ; [BRU2, 67]

Melitaea shandura Evans, 1924
Afghanistan, Pamirs, Pakistan, NW.India. See [About maps] Melitaea shandura Evans, 1924; J. Bombay nat. Hist. Soc. 30 (1): 90, TL: Shandur, Chitral [NW.India]
shandura Evans, 1912; (nom. nud.)
Melitaea shandura, NSG Voucher Specimen [Wahlberg]
Didymaeformia shandura ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea shandura ; [BRU2: 68, pl. 43, f. 31-33]
M. s. pavlitzkajana Sheljuzhko, 1943
W.Pamirs Melitaea shandura apsara Wyatt, 1961; J. Lep. Soc. 15 (1): 6, TL: above Kars Pass, Shiva Mts, N. Badakhshan, 12500-14000ft, Afghanistan
Melitaea shandura pavlitzkajana ; [BRU2, 68]

Melitaea athene Staudinger, 1881
Saur, Tarbagatai Melitaea athene Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 266, TL: Saisan [Saur Mts. near Zaisan, E.Kazakhstan]
Melitaea athene, NSG Voucher Specimen [Wahlberg]
Didymaeformia athene ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea athene ; [BRU2: 65, pl. 43, f. 34-36]

Melitaea trivia ([Schiffermüller], 1775)
-. See [About maps] Lesser Spotted Fritillary Desert Fritillary Braunlicher Scheckenfalter
Papilio trivia [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179, TL: Vienna
Papilio iphigenia Esper, 1782; Die Schmett. Th. I, Bd. 2 (5): 128, pl. 77, f. 1-2 (preocc.)
Papilio antigonus Herbst, 1800
cleo (Latreille, 1803)
Melitaea trivia interrupta Skala, 1907
Melitaea didyma aabaca Fruhstorfer, 1917
Melitaea trivia lathon Fruhstorfer, 1917
Melitaea catapeloides Stauder, 1918
Melitaea trivia tigris Higgins, 1941
Melitaea trivia, NSG Voucher Specimen [Wahlberg]
Melitaea trivia, Moths and Butterflies of Europe [MBE]
Melitaea trivia, Europaische Schmetterlinge [Mario Maier]
Melitaea trivia, Captain's European Butterfly Guide [Simon Coombes]
Didymaeformia trivia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea trivia ; [H&R, 118]; [BOW: pl. 3, f. 5]; [EBW]; [BRU2: 62, pl. 40, f. 1-36]; [Otakar Kudrna]

Larva on Verbascum thapsus [H&R]

M. t. fascelis (Esper, 1783)
S.Europe Papilio fascelis Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 171, pl. 88, f. 3-4
minor Ksienschopolsky, 1912
striata Ksienschopolsky, 1912
paula Obraztsov, 1936
Melitaeta trivia fascelis ; [BRU2, 62]

Larva on Scrophularia, Eremostachys [BRU2, 62]

M. t. uvarovi Gorbunov, 1995
Kazakhstan, SW.Siberia, Tarbagatai, Saur, Dzhungarsky Alatau Melitaea trivia uvarovi ; [BRU2, 62]
M. t. singularia Korshunov, 1995
Tuva Melitaea trivia singularia, Lepidoptera collection [SZMN]
Melitaea trivia singularia ; [BRU2, 62]
M. t. caucasi Verity, 1922
Caucasus, Transcausia Melitaea trivia caucasi ; [BRU2, 62]
M. t. nana Staudinger, 1871
Kopet-Dagh
pseudodidyma Rebel, 1905
syriaca Rebel, 1905
aabaca Fruhstorfer, 1917
wyatti Gross & Ebert, 1975
Melitaea trivia nana ; [BRU2, 62]
M. t. catapelia Staudinger, 1886
Ghissar, Alai, Tian-Shan Melitaea trivia bactriana Shchshetkin, 1984
= ; [BRU2, 62]
Melitaea trivia catapelia ; [BRU2, 62]
M. t. chorosana Shchetkin, 1984
S.Ghissar Melitaea trivia chorosana ; [BRU2, 62]
M. t. petri Shchetkin, 1984
Darvaz Melitaea trivia petri ; [BRU2, 62]
M. t. nativa Tuzov, 2000
W.Pamirs Melitaea trivia nativa Tuzov, 2000; [BRU2, 62 (repl.)]
Melitaea catapelia shugnana Shchetkin, 1984; (preocc. Melitaea didyma shugnana Sheljuzhko, 1929)
M. t. ignasiti Sagarra, 1926²
Portugal trivia ignasiti Sagarra, 1926²; , TL: Portugal
Melitaea trivia ignasiti ; [H&R, 118]

Melitaea robertsi Butler, 1880
Baluchistan, Chitral. See [About maps] Baluchi Fritillary
Melitaea robertsi Butler, 1880; Proc. zool. Soc. Lond. 1880 (3) : 406, pl. 39, f. 2
= Melitaea trivia ; [Wahlberg]

Melitaea yuenty Oberthür, 1888
W.China. See [About maps] Melitaea yuenty Oberthür, 1888; Etudes d'Ent. 11: 17, TL: Tatsienlu
Melitaea yuenty yuenty, Butterflies in Indo-China [Yutaka Inayoshi]
Didymaeformia yuenty ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea yuenty ; [BOW: pl. 196, f. 9]; [MRS, 587]
M. y. batangensis Belter, 1944
Melitaea yuenty batangensis Belter, 1944; , TL: Batang
Melitaea yuenty batangensis ; Huang, 2001

Melitaea lutko Evans, 1932
W.Himalayas. See [About maps] Melitaea lutko Evans, 1932; Indian Butterflies (edn. 2): 185
Melitaea lutko, NSG Voucher Specimen [Wahlberg]
Melitaea lutko ; [BOW: pl. 196, f. 4]
?Melitaea lukto[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name) [spell.?]

Melitaea mimetica Higgins, 1940
Pakistan, Afghanistan, Turkmenia. See [About maps] Melitaea mimetica Higgins, 1940; Entomologist 73: 52, TL: Baluchistan
= ; [Wahlberg]
Melitaea mimetica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [BRU2: 61, pl. 44, f. 10-12]

Melitaea romanovi Grum-Grshimailo, 1891
N.China, Mongolia, Transbaikalia. See [About maps] Melitaea romanovi Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4) : 454, TL: [Gansu, China]
Melitaea romanovi, NSG Voucher Specimen [Wahlberg]
Melitaea romanovi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [MRS, 587]; [BRU2: 65, pl. 44, f. 16-18]
N.China Melitaea romanovi romanovi ; [BRU2, 65]
M. r. puella Higgins, 1941
Transbaikalia Melitaea romanovi puella, Lepidoptera collection [SZMN]
Melitaea romanovi puella ; [Oleg Kosterin]; [BRU2, 65]
[puella-1fv.jpg] 800x600 (~103Kb) underside female The southern slope, covered with a short-grass steppe, of the hill Chikhalan on the northern bank of a brackish Lake Zun-Torei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 12th July 1996, Photo © Oleg Kosterin [puella-2f.jpg] 600x467 (~24Kb) female A short-grass steppe, on the northern bank of a brackish Lake Zun-Torei at the foot of the hill Chikhalan, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 13th July 1996 (captured by a spider), Photo © Oleg Kosterin

M. r. shanshiensis (Murayama, 1955)
N.China (Shansi) Euphidryas[sic] romanovi shanshiensis Murayama, 1955; Tyō to Ga 6 (1) : 1, f. 2, TL: Tai-yuan, Shansi, North China

Melitaea arduinna (Esper, 1783)
SEEU, S.Russia, Asia Minor, Iran, C.Asia. See [About maps] Freyer's Fritillary
Papilio arduinna Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 169, pl. 87, f. 4, TL: [Kamyshin, Volgograd Region, Russia]
Melitaea arduinna, NSG Voucher Specimen [Wahlberg]
Melitaea arduinna ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BOW: pl. 2, f. 28 (text only)]; [H&R, 114]; [BRU2: 72, pl. 45, f. 1-9]; [Otakar Kudrna]

Larva on Centaurea behen, (Transcaucasia) [BRU2, 72]

SEEU, SW.Siberia, Altai, Alai, N.Tian-Shan, Dzhungarsky Alatau, Tarbagatai, Saur Melitaea arduinna arduinna ; [BRU2, 72]
M. a. uralensis Eversmann, 1844
arduinna uralensis Eversmann, 1844; , TL: S.Urals
Melitaea arduinna uralensis ; [BRU2, 72]
M. a. rhodopensis Freyer, [1836]
Melitaea rhodopensis Freyer, [1836]; Neuere Beitr. Schmett. 3 : 3, pl. 193, f. 1, TL: ?Macedonia "europäischen Türkei"
rhodopensis ; Abadjiev, 2002, Neue Ent. Nachr. 23: 45
Melitaea arduinna rhodopensis ; [H&R, 114]
M. a. kocaki Wagener & Gross, 1976
Transcaucasis Melitaea arduinna kocaki ; [BRU2, 72]
M. a. evanescens Staudinger, 1886
Kopet-Dagh, Ghissar-Darvaz, W.Pamirs Melitaea arduinna evanescens ; [BRU2, 72]

Melitaea avinovi Sheljuzhko, 1914
Afghanistan, W.Pamirs. See [About maps] Melitaea avinovi Sheljuzhko, 1914; Dt. Ent. Z. Iris 28 (1) : 20, TL: [Nyut, Ishkashimsky Mts., W.Pamirs, Tajikistan]
Melitaea avinovi, NSG Voucher Specimen [Wahlberg]
Melitaea avinovi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BRU2: 74, pl. 44, f. 34-36]

Melitaea agar Oberthür, 1888
Tibet. See [About maps] Melitaea agar Oberthür, 1888; Etudes d'Ent. 11: 18
Melitaea agar Oberthür, 1886
Melitaea didyma wardi Watkins, 1927
Melitaea didyma baileyi Watkins, 1927
Melitaea agar ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BOW: pl. 196, f. 2]; [MRS, 587]
Melitaea agar baileyi ; [MRS, 587]

Melitaea sarvistana Wiltshire, 1941
Iran. See [About maps] Melitaea sarvistana Wiltshire, 1941; J. Bomb. Nat. Hist. Soc. 42 (3): 473
Melitaea sarvistana, NSG Voucher Specimen [Wahlberg]
Cinclidia sarvistana ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea sarvistana ; Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note)

Melitaea meherparvari Carbonell, 2007
Iran. See [About maps] Melitaea meherparvari Carbonell, 2007; Bull. Soc. ent. Fr. 112 (3): 334, TL: O. Iran, "ca. Mamulan, 900-950m, Lorestan

The "cinxia" species-group

fiMelitaea cinxia (Linnaeus, 1758)
EU, Morocco, Russia, W.Asia, Amurland. See [About maps] Täpläverkkoperhonen Glanville Fritillary Hökblomsternätfjäril Pricknätfjäril Wegerich-Scheckenfalter Le Damier Doncella punteada Przeplatka cinksia
Papilio cinxia Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 480, TL: Sweden
Papilio pilodellae Rottemburg, 1775
Papilio pilosella Rottemburg, 1775
Papilio delia Denis & Schiffermüller, 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179
Papilio abacus Rezius, 1783; (nom. nud.)
Papilio cinxia var. fulla Quensel, 1791
horvathi Abafi-Aigner, 1906
kasyi Gross & Ebert, 1975
Melitaea cinxia, NSG Voucher Specimen [Wahlberg]
Melitaea cinxia, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea cinxia, Svenska fjärilar [SVF]
Melitaea cinxia, Norges Sommerfugler [NORLEP]
Melitaea cinxia, Moths and Butterflies of Europe [MBE]
Melitaea cinxia, Europaische Schmetterlinge [Mario Maier]
Melitaea cinxia, Captain's European Butterfly Guide [Simon Coombes]
Melitaea cinxia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BOW: pl. 2, f. 28]; [EBW]; [H&R, 111]; [MRS, 587]; [BRU2: 71, pl. 44, f. 25-33]; [Otakar Kudrna]
[cinxia-1u.jpg] 415x463 (~38Kb) upperside Finland, A: Lemland, Flaka, 6/1995, Photo © Tero Piirainen [cinxia-2v.jpg] 533x400 (~33Kb) underside A clearing in a pine forest at the village Nizhnii Tsasuchei, Onon District, SE Chita Province (Dahuria), S Transbaikalia, Siberia, Russia. 2nd July 1996, Photo © Oleg Kosterin [cinxia-3l.jpg] 550x371 (~54Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
[cinxia-3u.jpg] 595x400 (~58Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch

Larva on Plantago lanceolata, P. media [SPRK], (in Finland, Estonia) Veronica spicata, (in C.Europe) V. teucrium [Marko Nieminen]


Morocco (Midle Atlas), W.Algeria Melitaea cinxia var. algirica Oberthür, 1915; Étud. Lépid. Comp. 10 : 427, pl. 288, f. 2339 (preocc. Rühl, 1892), TL: Sebdou (Algeria)
Melitaea cinxia eupompe Hemming, 1933; Entomologist 66: 276 (repl. algirica Oberthür, 1915)
Melitaea cinxia cinxia ; [BMAT: 51, pl. 16, f. 22-27]

Larva on Centaurea pullata, Scorzonera pygmaea [BMAT], (in Spain) Plantago spp., (in Israel) Veronica macrostachya [BMAT]


eupompe
[eupompe-1v.jpg] 600x435 (~30Kb) underside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre

M. c. clarissa Staudinger, 1901

Melitaea cinxia australis Verity, 1916
Melitaea cinxia narbonensis Fruhstorfer, 1917
Melitaea cinxia sicula Ragusa, 1920
Melitaea cinxia deva Hemming, 1932
Melitaea cinxia castiliana Turati, 1920
Melitaea cinxia carpetana Verity, 1929
M. c. atlantis Le Cerf, 1923
Morocco (High Atlas, Toubkal Massif) Melitaea cinxia atlantis Le Cerf, 1923; Bull. Soc. ent. Fr. 1923 (17) : 224, TL: Grand Atlas, Djebel Tachdirt, 3100-3250m
Melitaea cinxia atlantis ; [H&R, 111]; [BMAT: 52, pl. 16, f. 28-32]
[atlantis-1u.jpg] 600x435 (~47Kb) upperside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre [atlantis-2v.jpg] 600x435 (~28Kb) underside Morocco, Djebel Azourki, Haut Atlas, Photo © Jean Delacre

M. c. oasis Huang & Murayama, 1992
Xinjiang Melitaea cinxia oasis Huang & Murayama, 1992; Tyō to Ga 43 (1) : 7, f. 20, TL: Jiadengyu, Altai, 1380-1600m
Melitaea cinxia oasis ; [MRS, 587]
M. c. tschujaca Seitz, 1908
Altai - E.Siberia Melitaea cinxia tschujaca Seitz, 1908; , TL: Altai
Melitaea cinxia arelatia Fruhstorfer, 1910; Ent. Zs. 24 (26) : 144, TL: "Dauphine, Plateau d'Emparis bei La Grave"
Melitaea cinxia tschujaca ; [BRU2, 71]
[tschujaca-1mu.jpg] 698x500 (~48Kb) upperside male Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2300 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin

M. c. amardea Grum-Grshimailo, 1895
M. c. heynei Rühl, 1895
Alai Melitaea cinxia heynei Rühl, 1895; , TL: Alai
Melitaea cinxia heynei ; [BRU2, 71]
M. c. sacarina Fruhstorfer, 1917
cinxia sacarina Fruhstorfer, 1917; , TL: Saratov
Melitaea cinxia sacarina ; [BRU2, 71]
M. c. karavajevi Obraztsov, 1936
Ukraine cinxia karavajevi Obraztsov, 1936; , TL: Zhurovka, Kherson Region, Ukraine
Melitaea cinxia karavajevi ; [BRU2, 71]

Melitaea arcesia Bremer, 1861
S.Siberia (mountains), C.Yakutia, Magadan, Amurland, Primorye, Mongolia, N.China - C.China. See [About maps] Blackvein Fritillary
Melitaea arcesia Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 : 466, TL: ;Baical and Dahuria
Melitaea arcesia leechi Alphéraky, 1895
Melitaea arcesia irma Higgins, 1941
Melitaea baikalensis Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 : 465, pl. 1, f. 6
Melitaea arcesia, NSG Voucher Specimen [Wahlberg]
Melitaea arcesia ; Bremer, 1861, Bull. Acad. Imp. Sci. St. Petersb. 3 Mélanges biol. St.-Pétersb. 3 (5-6): 545
Melitaea baikalensis ; Kosterin
Melitaea arcesia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [BOW: pl. 196, f. 5]; [BRU2: 71, pl. 41, f. 4-9]
Transbaikalia Melitaea arcesia arcesia ; [BRU2, 71]
M. a. minor Elwes, 1899
Altai Melitaea baicalensis minor ;
Melitaea arcesia minor ; [BRU2, 71]
[minor-1fv.jpg] 1060x700 (~83Kb) underside female An alpine meadow in the valley of a headwater of the Chikty Rivulet, 2500 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin [minor-1mu.jpg] 460x528 (~42Kb) upperside male An alpine meadow on a southern slope in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin [minor-1mv.jpg] 672x652 (~74Kb) underside male An alpine meadow on a southern slope in the valley of the left headwater of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Myosotis sp.), Photo © Oleg Kosterin

M. a. chuana Grum-Grshimailo, 1893
Melitaea baicalensis var. chuana Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 (1-2): 128, TL: Amdo
Melitaea chuana, NSG Voucher Specimen [Wahlberg]
Melitaea chuana ;
Melitaea arcesia chuana ; [BRU2, 71]
[chuana-1fu.jpg] 1200x1263 (~294Kb) upperside female China, Qinghai prov., Caka nuo vill (3500m), 19.7.2010, Photo © A. Timchenko leg. [chuana-1mu.jpg] 1500x967 (~255Kb) upperside male China, Qinghai prov., Caka nuo vill (3500m), 19.7.2010, Photo © A. Timchenko leg. [chuana-1mv.jpg] 1500x1118 (~260Kb) underside male China, Qinghai prov., Caka nuo vill (3500m), 19.7.2010, Photo © A. Timchenko leg.

sikkimensis Moore, 1901; Lepidoptera Indica, 5 : 5
[sikkimensis-1fu.jpg] 1400x935 (~188Kb) upperside female China, S.Tibet, 10 km N. Nyalam town (4300m), 6.7.2010, Photo © A. Timchenko leg. [sikkimensis-1mu.jpg] 1400x1047 (~234Kb) upperside male China, S.Tibet, 10 km N. Nyalam town (4300m), 6.7.2010, Photo © A. Timchenko leg. [sikkimensis-1mv.jpg] 1400x1436 (~280Kb) underside male China, S.Tibet, 10 km N. Nyalam town (4300m), 6.7.2010, Photo © A. Timchenko leg.
[sikkimensis-1mx.jpg] 900x701 (~88Kb) male China, S.Tibet, 10 km N. Nyalam town (4300m), 6.7.2010, Photo © A. Timchenko leg.

M. a. carmana Fruhstorfer, 1915
Sayan Melitaea carmana Fruhstorfer, 1915
dabanica Baranchikov, 1979
Melitaea arcesia carmana ; [BRU2, 71]
M. a. rucephala Fruhstorfer, 1915
Inn-Shan Mts. Melitaea arcesia rucephala ; [BRU2, 71]

Melitaea sindura Moore, 1865
Kashmir, Chitral, Tibet. See [About maps] Melitaea sindura Moore, 1865; Proc. zool. Soc. Lond. 1865 (2) : 496
Melitaea sindura ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [BOW: pl. 195, f. 7]
Melitaea arcesia sindura ;
?M. s. honei Belter, 1942
Melitaea sindura honei Belter, 1942; , TL: NW.Yunnan

Melitaea balbita Moore, 1874
Pakistan, W.Himalayas. See [About maps] Melitaea balbita Moore, 1874; Proc. zool. Soc. Lond. 1874 (1) : 268
Melitaea balbita ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea arcesia balbita ;

Melitaea amoenula C. & R. Felder, [1867]
Tibet. See [About maps] Melitaea amoenula C. & R. Felder, [1867]; Reise Fregatte Novara, Bd 2 (Abth. 2) (3): 392
Melitaea amoenula, NSG Voucher Specimen [Wahlberg]
Melitaea amoenula ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea arcesia amoenula ;

Melitaea jezabel Oberthür, 1888
Tibet. See [About maps] Melitaea jezabel Oberthür, 1888; Etudes d'Ent. 11: 17
Melitaea jezabel Oberthür, ?1896
Melitaea jezabel yunnana Watkins, 1927; , TL: Shuantan, Loma valley, NW.Yunnan
= ; Huang, 2001, Neue Entomologische Nachrichten, 51: 103
Melitaea jezabel ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [MRS, 588]
Melitaea arcesia jezabel ;

Melitaea bellona Leech, [1892]
W.China. See [About maps] Melitaea bellona Leech, [1892]; Butts China Japan Corea (1): 219
Melitaea bellona, NSG Voucher Specimen [Wahlberg]
Melitaea bellona ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); [BOW: pl. 196, f. 3]; [MRS, 588]
Melitaea arcesia bellona ;

fiMelitaea diamina (Lang, 1789)
CEU, S.Europe, S.Siberia, NE.China, S.Ussuri, Korea, Japan. See [About maps] Tummaverkkoperhonen False-heath Fritillary Kovallnätfjäril Sotnätfjäril Silberscheckenfalter Doncella oscura Przeplatka diamina
Papilio diamina Lang, 1789; Verz. Schmett. Gegend. Augsburg (ed. 2) : 44, TL: Augsburg
Melitaea dictynna alpestris Fruhstorfer, 1917
Melitaea diamina, NSG Voucher Specimen [Wahlberg]
Melitaea diamina, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea diamina, Svenska fjärilar [SVF]
Melitaea diamina, Norges Sommerfugler [NORLEP]
Melitaea diamina, Moths and Butterflies of Europe [MBE]
Melitaea diamina, Europaische Schmetterlinge [Mario Maier]
Melitaea diamina, Captain's European Butterfly Guide [Simon Coombes]
Melitaea diamina ; [BOW: pl. 3, f. 1]; [EBW]; [H&R, 118]; [MRS, 588]; [BRU2: 75, pl. 47, f. 10-18]; [Otakar Kudrna]
[diamina-1u.jpg] 500x406 (~27Kb) upperside FINLAND: Orivesi, 1995, Photo © Juha Salokannel [diamina-2u.jpg] 500x412 (~32Kb) upperside Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen [diamina-3u.jpg] 500x419 (~26Kb) upperside Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen
[diamina-4.jpg] 500x595 (~64Kb) Finland, Orivesi, Siitama, 4.7.1996, Photo © Tero Piirainen [diamina-5fu.jpg] 965x591 (~125Kb) upperside female Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov [diamina-5mu.jpg] 775x668 (~104Kb) upperside male Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov

Larva on Valeriana officinalis, V. sambucifolia [SPRK], Veronica spp., Valeriana spp., Melampyrum spp. [MRS], Valeriana, Plantago, Veronica, Melampyrum, Patrinia, Polygonum [BRU2, 75]


Melitaea diamina diamina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
M. d. hebe (Borkhausen, 1793)
E.Europe - S.Siberia - Amur, Ussuri Papilio hebe Borkhausen, 1793
Papilio dictynna Esper, 1778; Die Schmett. Th. I, Bd. 1 (8): pl. 48, f. 2a, b
Papilio corythalia Hübner, [1799-1800]; Samml. eur. Schmett. [1] : pl. 3, f. 15-16
orthia Hübner, 1818
lebedevi Obraztsov, 1936
Mellicta dictynna ; [MRS, 586]
Cinclidia orthia ; Assmann, 1850, Zs. Ent. Breslau (Lepid.) (15): 42
Melitaea diamina hebe ; [BRU2, 75]
M. d. badukensis Alberti, 1969
Caucasus Major
M. d. vernetensis Rondou, 1902²
diamina vernetensis Rondou, 1902²; , TL: Vernet-les-Bain, E. Pyrenees
Melitaea diamina vernetensis ; [H&R, 119]
M. d. codinai Sagarra, 1932²
diamina codinai Sagarra, 1932²; , TL: Tortosa, NE.Spain
Melitaea diamina codinai ; [H&R, 119]
M. d. erycinides Staudinger, 1892
Melitaea diamina erycinides Staudinger, 1892; , TL: Kentei Mts.
Melitaea diamina erycinides ; [Oleg Kosterin]; [BRU2, 75 (?)]
[erycinides-1mu.jpg] 400x300 (~30Kb) upperside male A meadow at tiny aspen at a granite rocky ridge crest on a moderate elevation Adon-Chelon, at the mountain Tsagan-Obo, 10 km NNW of the village Tasyrkhoi, SE Chita region (Dahuria), Transbaikalia, Siberia, Russia. 9th July 1996, Photo © Oleg Kosterin

M. d. erycina (Lederer, 1853)
diamina erycina (Lederer, 1853); , TL: Altai
Mellicta dictynna erycina ; [MRS, 586]
Melitaea diamina erycina ; [BRU2, 75 (?)]

Melitaea protomedia Ménétriés, 1859
Amur, C.China, E.China, Korea, Japan. See [About maps] Melitaea protomedia Ménétriés, 1859; Bull. phys-math. Acad. Sci. St. Pétersb. 17 (14) : 214, TL: [Amur region, Russia]
Melitaea protomedia ab. argentea Fixsen, 1887; in Romanoff, Mém. Lépid. 3 : 301
Melitaea diamina protomedia ; [MRS, 588]
Melitaea protomedia ; [BOW: pl. 3, f. 1 (text only)]; [BRU2: 75, pl. 47, f. 19-21]

Larva on Veronica [BRU2, 76]

Amur, Ussuri Melitaea protomedia protomedia ; [BRU2, 75]
M. p. regama Fruhstorfer, 1915
SW.China - Korea protomedia regama Fruhstorfer, 1915; , TL: Chang Yang, SW.China
Melitaea diamina regama ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [MRS, 588]
Melitaea proomedia regama ; [BRU2, 75]

The "minerva" species-group
"minerva" ; Achtelik, 1999, Atalanta 30 (1/4): 63

Melitaea minerva Staudinger, 1881
Tian-Shan. See [About maps] Melitaea minerva Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 289, TL: Ala Tau [Dzhungarsky]
Melitaea minerva, NSG Voucher Specimen [Wahlberg]
Melitaea minerva ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BRU2: 70, pl. 42, f. 1-15]
N.Tian-Shan, Dzhungarsky Alatau Mts. Melitaea minerva minerva ; [BRU2, 70]
M. m. pseudotersa Kolesnichenko & Churkin, 2000
Inner Tian-Shan (Baidulu Mts.) Melitaea minerva pseudotersa Kolesnichenko & Churkin, 2000; , TL: Inner Tian-Shan (Baidulu Mts.)
Melitaea minerva pseudotersa ; [BRU2, 70]
M. m. repens Kolesnichenko & Churkin, 2000
Kara-Tau Melitaea minerva repens Kolesnichenko & Churkin, 2000; , TL: Kara-Tau Mts.
Melitaea minerva repens ; [BRU2, 70]
M. m. tersa Kolesnichenko & Churkin, 2000
Karzhantau, Chatkal Melitaea minerva tersa ; [BRU2, 70]
M. m. acerba Kolesnichenko & Churkin, 2000
Chatkalsky Mts. (Lake Sary-Chelek) Melitaea minerva acerba Kolesnichenko & Churkin, 2000; , TL: Chatkalsky Mts. (Lake Sary-Chelek)
Melitaea minerva acerba ; [BRU2, 70]

Melitaea pallas Staudinger, 1886
-. See [About maps] Melitaea pallas Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 234, TL: [N. slope of the Alai Mts., Kirghizia]
Melitaea pallas, NSG Voucher Specimen [Wahlberg]
Melitaea pallas ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BOW: pl. 196, f. 6]; [MRS, 587]; [BRU2: 70, pl. 42, f. 16-30]
Alai, Transalai, Inner Tian-Shan Melitaea pallas pallas ; [BRU2, 70]
M. p. iracunda Kolesnichenko & Churkin, 2000
Talassky Ala-Tau, Chatkalsky Mts. Melitaea pallas iracunda ; [BRU2, 70]
M. p. mendax Kolesnichenko & Churkin, 2000
Kirghizsky Mts. (Alamedin River) Melitaea pallas mendax ; [BRU2, 71]
M. p. interpres Kolesnichenko & Churkin, 2000
Naryn, Baidulu Mts. Melitaea pallas interpres ; [BRU2, 71]
M. p. vaschenkoi Kolesnichenko & Churkin, 2000
E.At-Bashi Mts. Melitaea pallas vaschenkoi ; [BRU2, 71]
M. p. pseudoalpina Kolesnichenko & Churkin, 2000
E.Pamirs (Sarukolsky Mts.) Melitaea pallas pseudoalpina ; [BRU2, 71]
M. p. koitezek Kolesnichenko & Churkin, 2000
W.Pamirs (Pass Koitezek) Melitaea pallas koitezek ; [BRU2, 71]

Melitaea turanica Erschoff, 1874
Darvaz Melitaea turanica Erschoff, 1874; in Fedschenko, Travels in Turkestan. 2 (5) : 15, pl. 1, f. 11-12, TL: Tajikistan, Darvaz, Petr I Mts., Ganishou loc. 1,900m
Melitaea pamira Staudinger, 1887; Stettin Ent. Ztg 48 (1-3) : 53
Melitaea expressa Grum-Grshimailo, 1887
Melitaea turanica, NSG Voucher Specimen [Wahlberg]
Melitaea turanica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); [BRU2: 69, pl. 41, f. 11-13]

Melitaea asteroida Staudinger, 1881
Dzhungarsky Alatau Melitaea asteroida Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 292, TL: Ala Tau [Dzhungarsky Alatau]
Melitaea asteroida, NSG Voucher Specimen [Wahlberg]
Melitaea asteroidea[sic] ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name)
Melitaea asteroida ; [BRU2: 68,pl. 41, . 13-15, pl. 86, f. 15-18 (missp.?)]
M. a. clara Staudinger, 1887
Melitaea clara Staudinger, 1887
M. a. uitasica Wagner, 1913
Melitaea uitasica Wagner, 1913

Melitaea ludmilla Churkin, Kolesnichenko & Tuzov, 2000
N.Tian-Shan, W.Tian-Shan. See [About maps] Melitaea ludmilla Churkin, Kolesnichenko & Tuzov, 2000; Helios 1: (61-87)
Melitaea ludmilla, NSG Voucher Specimen [Wahlberg]
Melitaea ludmilla ; [BRU2: 68, pl. 41, f. 28-33]
Kirghizsky Mts. Melitaea ludmilla ludmilla ; [BRU2, 68]
M. l. serene Tuzov, 2000
W.Tian-Shan Melitaea ludmilla serene ; [BRU2, 68]

Melitaea solona Alphéraky, 1881
-. See [About maps] Melitaea solona Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 404, TL: [Kulja, Kunges Valley, W.China]
Melitaea solona, NSG Voucher Specimen [Wahlberg]
Melitaea solona ; [BRU2: 69, pl. 41, f. 16-27, pl. 86, f. 19-22]
E.Tian-Shan, W.Tibet? Melitaea solona solona ; [BRU2, 69]
M. s. filipjevi Churkin, Kolesnichenko & Tuzov, 2000
N.Tian-Shan (Zailiisky Alatau Mts.) Melitaea solona filipjevi Churkin, Kolesnichenko & Tuzov, 2000; Helios 1: (61-87)
Melitaea solona filipjevi ; [BRU2, 69]
M. s. plyushchi Churkin, Kolesnichenko & Tuzov, 2000
C.Tian-Shan (Sary-Dzhaz, Kokshaal-Tau) Melitaea solona plyushchi Churkin, Kolesnichenko & Tuzov, 2000; Helios 1: (61-87)
Melitaea solona plyushchi ; [BRU2, 69]
M. s. pletnevi Churkin, Kolesnichenko & Tuzov, 2000
E.Alai, Inner Tian-Shan Melitaea solona pletnevi Churkin, Kolesnichenko & Tuzov, 2000; Helios 1: (61-87)
Melitaea solona pletnevi ; [BRU2, 69]
M. s. evadne Hemming, 1934
E.Transalai solona evadne Hemming, 1934; Stylops 3 : 98
pallida Staudinger, 1901; (preocc.)
Melitaea solona evadne ; [BRU2, 69]

Melitaea sultanensis Staudinger, 1886
-. See [About maps] Melitaea parthenie var. sultanensis Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 234, TL: Samarkand (region) [Ghissarky Mts., Uzbekistan]
Melitaea sultanensis, NSG Voucher Specimen [Wahlberg]
Melitaea minerva sultanensis ; Higgins, 1940, Entomologist 73
Melitaea sultanensis ; Achtelik, 1999, Atalanta 30 (1/4): 77, pl. 3, f. 1-2, pl. 4, f. 1-2, pl. 5; [BRU2: 69, pl. 41, f. 34-39]
Ghissar Melitaea fumarata Achtelik, 1999; Atalanta 30 (1/4): 75, pl. 1, f. 5-6, pl. 2, f. 5-6, TL: Tadschikistan, Hisarskij Ch., Pass Anzob, 3500m
= ; [BRU2, 69]
Melitaea sultanensis sultanensis ; [BRU2, 69]
fumurata[sic, recte fumarata] ; [BRU2, 69]
M. s. gabrielae Achtelik, 1999
Darvaz Melitaea gabrielae Achtelik, 1999; Atalanta 30 (1/4): 73, pl. 1, f. 7-8, pl. 2, f. 7-8, TL: Tadzikistan, Darvar-Chorzatvschah
Melitaea sultanensis gabrielae ; [BRU2, 69]
M. s. sebastiani Achtelik, 1999
Alai, Transalai Melitaea sebastiani Achtelik, 1999; Atalanta 30 (1/4): 67, pl. 1, f. 1-2, pl. 2, f. 3-4, TL: Kirghisia, Mt. Alai, Isfara-Fl., 4000m
Melitaea sultanensis sebastiani ; [BRU2, 69]
M. s. palamedes Grum-Grshimailo, 1890
Tian-Shan, E.Alai Melitaea palamedes Grum-Grshimailo, 1890; in Romanoff, Mém. Lép. 4 : 436, pl. 10, f. 7
Melitaea danieli Achtelik, 1999; Atalanta 30 (1/4): 66, pl. 1, f. 1-2, pl. 2, f. 1-2, TL: Alma Ata, Medeo, 2200m
= ; [BRU2, 70]
Melitaea minerva palamedes ;
Melitaea sultanensis palamedes ; [BRU2, 70]
M. s. balbina Tytler, 1926
N.Chitral Melitaea balba balbina Tytler, 1926; J. Bombay nat. Hist. Soc. 31 (3): 583, TL: Shandur Pass between Gilgit and Chitral, 12200ft
Melitaea minerva balbina ; Higgins, 1940, Entomologist 73; Tshikolovets, 1997, The Butterflies of Pamir: ?
Melitaea balbina ; Achtelik, 1999, Atalanta 30 (1/4): 80, pl. 3, f. 7-8, pl. 4, f. 7-8
Melitaea sultanensis balbina ; [BRU2, 70]

Melitaea elisabethae Avinoff, 1910
Pamir, Badakhshan, Hindu-Kush, Tajikistan, Afghanistan. See [About maps] Melitaea elisabethae Avinoff, 1910; Horae Soc. Ent. Ross. 39 : 235, pl. 14, f. 6-7, TL: south from Alitshur, S. Pamir
Melitaea minerva elizabethae[sic] ; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7); D'Abrera, 1992, Butterflies of the Holarctic Region, Part II
Melitaea elisabethae ; Achtelik, 1999, Atalanta 30 (1/4): 82, pl. 3, f. 3-4, pl. 4, f. 3-4
?Melitaea sultanensis elisabethae ; [BRU2, 70 (?)]

Melitaea balba Evans, 1912
W.Himalayas, Karakorams. See [About maps] Melitaea balbita r. balba Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 582, TL: Chitral
Melitaea arcesia balba ; Evans, 1927, Indian Butterflies (ed. 1): 121
Melitaea minerva balba ; Evans, 1932, Indian Butterflies (edn. 2): 186; Higgins, 1941, Trans. ent. Soc. Lond. 91 (7):
Melitaea balba ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 165 (name); Achtelik, 1999, Atalanta 30 (1/4): 79, pl. 3, f. 5-6, pl. 4, f. 5-6

The "phoebe" species-group

Melitaea phoebe (Goeze, [1779])
CEU, SEU, tr, NAF, C.Asia, China. See [About maps] Knapweed Fritillary Flockenblumenfalter Le Grand Damier Doncella mayor Przeplatka febe
Papilio phoebe Goeze, [1779]
Papilio phoebe [Schiffermüller], 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 179 (nom. nud.), TL: Vienna
Papilio paedotrophos Bergsträsser, 1780; Nomen. Ins., 4: 14
Papilio corythallia Esper, 1781; Die Schmett. Th. I, Bd. 2 (2): 67, (4): pl. 72, f. 2
phoebe (Knoch, 1783); (preocc.)
Papilio tremulae Piller & Miterpacher, 1783
f. melanina Bonaparte, 1831; (dark)
f. aetherea Eversmann, 1851; (light)
Melitaea phoebe ab. tatara Krulikovsky, 1890; Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 236, pl. 8, f. g
Melitaea phoebe aethereaeformis Verity, 1919
Melitaea phoebe alternans Seitz, 1909
Melitaea phoebe canellina Stauder, 1922
Melitaea phoebe dorae Graves, 1925
Melitaea phoebe gerinia Fruhstorfer, 1916
Melitaea phoebe koios Fruhstorfer, 1908
Melitaea phoebe minoa Fruhstorfer, 1917
Melitaea phoebe ogygia Fruhstorfer, 1908
Melitaea phoebe rovia Fruhstorfer, 1919
Melitaea phoebe tusca Verity, 1919
Melitaea phoebe, NSG Voucher Specimen [Wahlberg]
Melitaea phoebe, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Melitaea phoebe, Moths and Butterflies of Europe [MBE]
Melitaea phoebe, Europaische Schmetterlinge [Mario Maier]
Melitaea phoebe, Captain's European Butterfly Guide [Simon Coombes]
Cinclidia phoebe ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea phoebe ; [BOW: pl. 3, f. 4, pl. 104, f. 16]; [EBW]; [H&R, 114]; [BOW: pl. 195, f. 6]; [Otakar Kudrna]
Melitaea (phoebe) phoebe ; [BRU2: 72, pl. 46, f. 12-14, pl. 47, f. 1-9]
[phoebe-1u.jpg] 951x624 (~151Kb) upperside Quinson/Verdon, Var, France, 8-89, Photo © S. Shuichi Haupt [phoebe-2l.jpg] 400x598 (~63Kb) larva Central Europe 1990-2000, Photo © Michael Borsch [phoebe-2p.jpg] 400x598 (~48Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch
[phoebe-2u.jpg] 598x400 (~78Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch [phoebe.jpg] 500x377 (~54Kb) Turunc, SW Turkey 18/04/95, Photo © Simon Coombes

Larva on Centaurea, less Plantago [H&R]


S.Europe, N.Caucasus, SW.Siberia, Altai tartara Krulikovsky, 1891
Melitaea phoebe var. ornata Christoph, 1893; Dt. Ent. Zeit. Iris, 6 (1) : 87
parva Caradja, 1895
uralensis Krulikovsky, 1897; (preocc.)
ufensis Krulikovsky, 1902
Melitaea phoebe lokris Fruhstorfer, 1908
sextilis Jachontov, 1908
Melitaea phoebe phoebe ; [BRU2, 72]
M. p. ottonis Fruhstorfer, 1916
Transcaucasia Melitaea phoebe caucasicola Verity, 1919
Melitaea phoebe ottonis ; [BRU2, 73]
M. p. enoch Higgins, 1941
Kopet-Dagh? Melitaea phoebe enoch Higgins, 1941; , TL: vicinity of Ashkhabad
Melitaea phoebe enoch ; [BRU2, 73]
M. p. wagneri Wnukowsky, 1929
Tian-Shan, Saur, Tarbagatai Melitaea phoebe alatauica Wagner, 1913; (preocc.)
Melitaea phoebe wagneri ; [BRU2, 73]
M. p. tungana Seitz, [1909]
S.Siberia (mountains) Melitaea phoebe tungana ; [BRU2, 73]
[tungana-1fuv.jpg] 700x821 (~120Kb) upperside underside female Russia: C.Tuva, W of Ujukskiy Mts., Kamennyi R. (1000m), 11-20.7.2003, Vashchenko S. N. leg. Photo © D. Smirnov

M. p. tungusa Herz, 1898
Yakutia Melitaea phoebe tungusa ; [BRU2, 73]
M. p. mandarina Seitz, [1909]
Amur, Mongolia, Transbaikalia Melitaea phoebe mandarina ; [BRU2, 73]
M. p. occitanica Staudinger, 1861
SWEU, Algeria, Morocco Melitaea phoebe v. occitanica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 8, TL: Spain
Melitaea phoebe ab. punica-powelli Oberthür, 1915; Étud. Lépid. Comp. 10 : pl. 288, f. 2338
Melitaea phoebe leechi Rothschild, 1917; Novit. Zool. 24 (1): 99 (preocc.), TL: Mogador [Essaouira] (Morocco)
Melitaea phoebe f. gaisericus Hemming, 1941; Proc. R. ent. Soc. Lond. 10 : 207 (repl. Melitaea phoebe leechi Rothschild)
Melitaea phoebe occitanica ; [BMAT: 52, pl. 17, f. 1-10]; [H&R, 115]

Larva on Rhaponticum acaule, Centaurea sp., (in Spain) Centaurea scabiosa, C. jacea, C. montana [BMAT]


gaisericus
[gaisericus-1u.jpg] 800x580 (~56Kb) upperside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [gaisericus-2u.jpg] 600x435 (~47Kb) upperside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [gaisericus-3v.jpg] 800x580 (~54Kb) underside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre

M. p. changaica Seitz, [1909]
Melitaea phoebe changaica Seitz, [1909]; , TL: Khangai Mts.
Melitaea phoebe changaica ; [Oleg Kosterin]; [BRU2, 73]
[changaica-1l.jpg] 600x800 (~115Kb) larva A meadow steppe on a moderate elevation Adon-Chelon at the mountain Tsagan-Obo, 10 km NNW of the village Tasyrkhoi, SE Chita region (Dahuria), Transbaikalia, Siberia, Russia. 19th June 1995 (mature larva), Photo © Oleg Kosterin [changaica-1p.jpg] 400x533 (~49Kb) pupa 3rd July, the caterpillar was collected on a meadow steppe on a moderate elevation Adon-Chelon at the mountain Tsagan-Obo, 10 km NNW of the village Tasyrkhoi, SE Chita region (Dahuria), Transbaikalia, Siberia, Russia. 19th June 1995, Photo © Oleg Kosterin [changaica-2mfv.jpg] 600x800 (~110Kb) underside male female The southern slope, covered with a short-grass steppe, of the hill Chikhalan on the northern bank of a brackish Lake Zun-Torei, Onon district, Chita Province, S Transbaikalia (Dahuria), Siberia, Russia. 12th July 1996, Photo © Oleg Kosterin

M. p. saturata Staudinger, 1892
Melitaea phoebe saturata Staudinger, 1892; , TL: Kentei Mts.
Melitaea phoebe saturata ; [Oleg Kosterin]; [BRU2, 73]
[saturata-1v.jpg] 400x437 (~38Kb) underside A meadow steppe on the Shipunikha Rivulet right bank terrace 2 km NE of the village Evsino, Iskitim District, Novosibirsk Province, West Siberia, Russia. 21st June 1997 (On Campanula sibirica L.), Photo © Oleg Kosterin


Melitaea punica Oberthür, 1876
-. See [About maps] Melitaea phoebe var. punica Oberthür, 1876; Étud. d'Ent. 1 : 25, pl. 1, f. 3, TL: Lambessa (Algeria)
= ?Melitaea phoebe occitanica Staudinger, 1861; [BMAT, 52]
Melitaea phoebe punica ; [H&R, 115]
Melitaea (phoebe) punica ; [BRU2: 73, pl. 46, f. 7-9, 15-17]
M. p. amanica Rebel, 1917
Transcaucasia Melitaea phoebe amanica Rebel, 1917
Melitaea punica amanica ; [BRU2, 73]
?M. p. telona Fruhstorfer, 1908
Israel, Jordan, Lebanon, Syria, Iraq, Iran, Transcaucasia, Anatolia, Balkan Peninsula Melitaea phoebe telona Fruhstorfer, 1908; Int. ent. Zs. 1 (41) : 310, TL: Greece, Poros

Melitaea pseudosibina Alberti, 1969
Caucasus, Transcaucasia. See [About maps] Melitaea pseudosibina Alberti, 1969; Dt. Ent. Z., N.F, 16: 192, pl. 1, f. 1c, 2c, TL: Itkol im Elbrus-Gebiet, 2100m
Melitaea (phoebe) pseudosibina ; [BRU2: 73, pl. 46, f. 1-3]

Melitaea aetherie Hübner, [1826]
NAF, Sicily, Iberian Peninsula. See [About maps] Aetherie Fritillary Doncella gaditana
Melitaea aetherie Hübner, [1826]; Samml. eur. Schmett. [1] : pl. 177, f. 875-878, TL: S.Spain
Melitaea aetherie, NSG Voucher Specimen [Wahlberg]
Melitaea aetherie, Moths and Butterflies of Europe [MBE]
Cinclidia aetherie ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea aetherie ; [H&R, 115]; [Otakar Kudrna]
M. a. algirica Rühl, 1892
Algeria, Morocco (Middle Atlas, Toubkal Massif), W.Tunisia (Kroumerie) Melitaea aetheria var. algirica Rühl, 1892; Palaearktischen Grosschmetterlinge: 389, TL: Algeria
Melitaea aetherie algirica ; [H&R, 115]; [BMAT: 53, pl. 17, f. 11-21]
[algirica-1u.jpg] 800x580 (~50Kb) upperside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [algirica-2u.jpg] 800x580 (~60Kb) upperside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [algirica-4v.jpg] 800x580 (~44Kb) underside Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre
[algirica-5fu.jpg] 600x435 (~80Kb) upperside female Morocco, Tizi-n-Tretten, Moyen Atlas, Photo © Jean Delacre [algirica-6u.jpg] 600x435 (~49Kb) upperside Morocco, Ito, Moyen Atlas , Photo © Michel Tarrier [algirica-7v.jpg] 600x435 (~27Kb) underside Morocco, Tizi-n-Tretten, Moyen Atlas , Photo © Michel Tarrier

Larva on Centaurea spp. [BMAT]



Melitaea sibina Alphéraky, 1881
Ghissa-Darvaz, Tian-Shan. See [About maps] Melitaea sibina Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 400, TL: Kouldja, Ili Valley
Melitaea sibina, NSG Voucher Specimen [Wahlberg]
Cinclidia sibina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea sibina ; [BOW: pl. 196, f. 8]; [BRU2: 74, f. 10-21]
Ili Valley Melitaea sibina sibina ; [BRU2, 74]
M. s. aulica Seitz, [1909]
Ghissar-Darvaz, Tianshan rama Higgins, 1941
Melitaea sibina aulica ; [BRU2, 74]
M. s. dschungarica Grum-Grshimailo, 1895
Dzhungarsky Alatau Melitaea sibina dschungarica ; [BRU2, 74]

Melitaea scotosia Butler, 1878
NE.China, Korea, Ussuri, Japan. See [About maps] Melitaea scotosia Butler, 1878; Cist. Ent. 2 (19) : 282, TL: Tokyo, Japan
yagei Nire, 1917
parascotosia Collier, 1933
Melitaea scotosia weiwueria Huang & Murayama, 1992; Tyō to Ga 43 (1) : 7, f. 19, TL: Jiadengyu, Altai, 1400m
Melitaea scotosia, NSG Voucher Specimen [Wahlberg]
Cinclidia scotosia ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea scotosia ; [MRS, 589]; [BRU2: 74, pl. 46, f. 4-6, 10-11]

Larva on Serratula coronata, Cirsium japonicum, Saussurea pulchella, Synurus excelsus [MRS]

M. s. butleri Higgins, 1940
Melitaea scotosia butleri Higgins, 1940; Entomologist 73: 53
= Melitaea scotosia ; [BRU2, 74]
Melitaea scotosia butleri ; [MRS, 589]

Melitaea collina Lederer, 1861
Middle East, Asia Minor. See [About maps] Melitaea collina Lederer, 1861; Wien. ent. Monats. 5 : 148, pl. 1, f. 4-5
Melitaea collina, NSG Voucher Specimen [Wahlberg]
Didymaeformia collina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)

Melitaea turkmanica Higgins, 1940
NE.Turkey, Armenia, Kopet-Dagh, Iran?. See [About maps] Melitaea turkmanica Higgins, 1940; Entomologist 73: 51, TL: "Askhabad" [Kopet-Dagh, Turkmenia]
Melitaea turkmanica, Lepidoptera collection [SZMN]
Didymaeformia turkmanica ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea turkmanica ; [BRU2: 74, pl. 44, f. 19-24]
Kopet-Dagh Melitaea turkmanica turkmanica ; [BRU2, 74]
M. t. vedica Nekrutenko, 1975
Armenia Melitaea vedica Nekrutenko, 1975; J. Lep. Soc. 29 (2): 102
Melitaea turkmanica vedica ; [BRU2, 74]

Melitaea consulis Wiltshire, 1941
Iran. See [About maps] Melitaea consulis Wiltshire, 1941; J. Bomb. Nat. Hist. Soc. 42: 9?
Melitaea consulis, NSG Voucher Specimen [Wahlberg]
Didymaeformia consulis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)

Melitaea kuchi Wyatt, 1961
Afghanistan. See [About maps] Melitaea kuchi Wyatt, 1961; J. Lep. Soc. 15 (1): 7, f. (m.gen), TL: Band-i-Amir, Hazarajt, N of Koh-i-Baba Mts., 11500ft
Didymaeformia kuchi ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)

Melitaea gina Higgins, 1941
Iran. See [About maps] Melitaea gina Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 353
Melitaea gina, NSG Voucher Specimen [Wahlberg]
Didymaeformia gina ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name)
Melitaea gina ; Weiss & Major, 2000, Atalanta 31 (1/2): 126 (note), pl. V, f. 9-12

The "miscellaneous?" species-group

Melitaea wiltshirei Higgins, 1941
Iran. See [About maps] Melitaea wiltshirei Higgins, 1941; Trans. ent. Soc. Lond. 91 (7): 237
Melitaea wiltshirei, NSG Voucher Specimen [Wahlberg]

Melitaea tangigharuensis de Freina, 1980
Afghanistan. See [About maps] Melitaea tangigharuensis de Freina, 1980; Atalanta 11 (1): (11-18)
Melitaea tangigharuensis ; Higgins, 1981, Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 166 (name); Carbonell, 2007, Bull. Soc. ent. Fr. 112 (3): 334 (note)

Melitaea paludani Clench & Shoumatoff, 1956
Afghanistan. See [About maps] Melitaea paludani Clench & Shoumatoff, 1956; Vidensk. Medd. Dansk Naturh. Forening 118: (141-192) , TL: Marak, 4500m, Afghanistan


21.9.2013 (70)

References:
[A. Timchenko]
Andrew Timchenko; Beijing, China (ex-Moscow, Russia)
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[AfrL]
Mark C. Williams; Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BAFR]
D'Abrera, 1980; Butterflies of the Afrotropical Region
[BHL]
The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[BMAT]
Tennent, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9
[BOW]
Lewis, 1974; Butterflies of the World; ISBN 0 245 52097 X
[BRU2]
Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2
[BnF]
Gallica / Bibliothčque nationale de France; Annales de la Société entomologique de France; Bulletin de la Sociéte entomologique de France
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[EBW]
Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[H&R]
Higgins, L.G, Riley, N.D, 1970; A Field Guide to the Butterflies of Britain and Europe
[HBN]
Francis Hemming, 1937; HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Jean Delacre]
http://homepage.mac.com/jdelacre
[Kimmo Silvonen]
Espoo, Finland; Kimmo's Lep Site
[LAMAS-2005]
Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MRS]
Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Marko Nieminen]
Department of Ecology and Systematics; Division of Population Biology; P.O. Box 17 (Arkadiankatu 7); FIN-00014 University of Helsinki; tel. +358-9-191 28779; fax. +358-9-191 28701; http://www.helsinki.fi/science/metapop/
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NBC]
Netherlands Centre for Biodiversity Naturalis; NBC naturalis
[NORLEP]
Norges Sommerfugler
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[S. Shuichi Haupt]
e-mail: S. Shuichi Haupt; http://www.tu-berlin.de/fb7/ioeb/biologie/stefan.htm
[SPRK]
Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN]
Lepidoptera; Siberian Zoological Museum
[Simon Coombes]
1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen]
Tampere, Finland; e-mail: Tero Piirainen
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[²]
This may require parentheses or not. I don't have the necessary information for this taxon.


Some related literature:
Abadjiev, 2002
Types of Balkan butterflies in the collection of The Natural History Museum, London Neue Ent. Nachr. 23: 3-53
Achtelik, 1999
Vier neue Arten aus der "Melitaea sultantensis-Gruppe" (Staudinger, 1886) Atalanta 30 (1/4): 63-86
Alberti, 1969
Neue oder bemerkenswerte Lepidoptera-formen aus den Grossen Kaukasus Dt. Ent. Z., N.F, 16: 189-203, 3pls
Alphéraky, 1881
Lépidoptčres du district de Kouldją et des montagnes environnantes. (1-3) Horae Soc. ent. Ross. 16 (3-4): 334-435, pl. 14-15 (1881), 17 (1-2): 15-103, pl. 1-3 (1882), 17 (3-4): 156-227, pl. 8-9 (1883)
Assmann, 1850
Eine Exkursion nach klarenkranst, und Die Raupe und Puppe der Melitaea Britomartis Ass. Zs. Ent. Breslau (Lepid.) (15): 37-42
Avinoff, 1910
[Contribution ą la faune des Rhopaloceres du Pamir] (in Russian) Horae Soc. Ent. Ross. 39 : 225-246, pl. 14
Bergsträsser, 1780
Nomenclatur und Beschreibung der Insecten in der Graftschaft Hanau-Münzenburg Nomen. Ins., 4: 47pp, pl. 73-96
Bremer, 1861
Neue Lepidopteren aus Ost-Sibirien und dem Amur-Lande gesammelt von Radde und Maack, bescrieben von Otto Bremer Bull. Acad. Imp. Sci. St. Petersb. 3 : 461-496, Mélanges biol. St.-Pétersb. 3 (5-6): 538-589
Bryk, 1940
Geographische Variabilität von Melitaea didyma (Esper) Folia Zoologica et Hydrobiologica 10 (2): 293-353, pl. 1-6
Butler, 1878
On some Butterflies recently sent home from Japan by Mr. Montagne Fenton Cist. Ent. 2 (19) : 281-286
Butler, 1880
On a collection of Lepidoptera from Candahar Proc. zool. Soc. Lond. 1880 (3) : 403-415, pl. 39
Carbonell, 2007
Melitaea meherparvari n. sp. du sud-ouest de l'Iran (Lep., Nymphalidae) Bull. Soc. ent. Fr. 112 (3): 334
Le Cerf, 1923
Lépidoptčres nouveaux du Maroc Bull. Soc. ent. Fr. 1923 (17) : 224
Christoph, 1893
Lepidoptera Nova Faunae Palaearcticae Dt. Ent. Zeit. Iris, 6 (1) : 86-96
Churkin, 2000
Taxonomical notes on Melitaea ambrisia Higgins, 1935 and Melitaea alraschid Higgins, 1941 stat. nov. from Central Asia Atalanta 31 (1/2): 113-122
Churkin, Kolesnichenko & Tuzov, 2000
Revision of the Melitaea asteroida species group (Lepidoptera, Nymphalidae) with new taxa descriptions Helios 1: 61-87
Churkin & Tuzov, 2000
Revision of Melitaea fergana Staudinger, 1882 (Lepidoptera, Nymphalidae) Helios 1: 33-60
Clench & Shoumatoff, 1956
Lepidoptera Rhopalocera (Insecta) from Afghanistan. The 3rd Danish Expedition to Central Asia (Zoological Results 21) Vidensk. Medd. Dansk Naturh. Forening 118 : 141-192, 23 fgs
D'Abrera, 1992
Butterflies of the Holarctic Region, Part II, Satyridae (concl.) & Nymphalidae (partim) Butterflies of the Holarctic Region, Part II
Eisner, 1942
Einige ergebnisse der Sichtung der Gattungen Melitaea und Argynnis im Rijksmuseum van Natuurlijke Historie, Leiden Zool. Meded. Leiden 24 (4): 116-124
Esper, 1778
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Band 1 Die Schmett. Th. I, Bd. 1 (1): 1-40, pl. 1-6 (1776), (2): 41-72, pl. 7-12 (1776), (3): 73-104, pl. 13-18 (1777), (4): 105-120, pl. 19-24 (1777), (5): 121-144, pl. 25-30 (1777), (6): 145-176, pl. 31-36 (1777), (7): 177-200, pl. 37-46 (1778), (8): 201-216, pl. 43-48 (1778), (9): 217-388, pl. 49-50 (1779)
Esper, 1781; Esper, 1782; Esper, 1783
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil I. Die Tagschmetterlinge. Fortsetzung. Band 2 Die Schmett. Th. I, Bd. 2 (1): 1-36, pl. 51-56 (1780), (2): 37-68, pl. 57-62 (1781), (3): 69-100, pl. 63-68 (1781), (4): 101-124, pl. 69-74 (1781), (5): 125-140, pl. 75-80 (1782), (7): 141-156, pl. 81-82 (1783), (8): 157-172, pl. 83-88 (1783), (9): 173-184, pl. 89-92 (1784), (28): 185-190, pl. 93 (1786)
Erschoff, 1874
Travels in Turkestan. Volume 2. Zoogeographical Investigations. Lepidoptera [in Russian] in Fedschenko, Travels in Turkestan. 2 (5) : 1-128, pl. 1-6
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1924
The identification of Indian Butterflies (5-8) J. Bombay nat. Hist. Soc. 30 (1): 72-96, (2): 322-351, pl. 26-27, (3): 610-639, pl. 28, (4): 726-776, pl. 29
Evans, 1932
The Identification of Indian Butterflies (edn. 2) Indian Butterflies (edn. 2): x + 454pp, 9 figs, 32 pls
Eversmann, 1847
Lepidoptera quaedam nova Rossiae et Sibiriae indigena descripsit et delineavit Bull. Soc. Imp. Nat. Moscou 20 (3) : 66-83, 6 pls.
Fabricius, 1807
Systema Glossatorum. Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Magazin f. Insektenk. (Illiger) 6: 277-295
C. & R. Felder, [1867]
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera Reise Fregatte Novara, Bd 2 (Abth. 2) (1): 1-136, pl. 1-21 (Rhop.) (C. & R. Felder, [1865]), (2): 137-378, pl. 22-47 (Rhop.) (C. & R. Felder, 1865), (3): 379-536, pl. 48-74 (Rhop.) (C. & R. Felder, 1867), (4): 537-548, pl. 75-120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) , (5): pl. 121-140, (erklärung) 1-10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Fixsen, 1887
Lepidoptera aus Korea in Romanoff, Mém. Lépid. 3 : 233-356, pl. 13-15
de Freina, 1980
Melitaea tangigharuensis n. sp., eine neue Nymphalidenart für Afghanistan nebst vergleichender Betraschtung zur M. sarvostama Wiltshire, 1941 Atalanta 11 (1): 11-18
Freyer, [1836]
Neuere Beiträge zur Schmetterlingskunde mit Abbildungen nach der Natur. (33-48) Neuere Beitr. Schmett. 3 : 1-20 (1836), : 21-52 (1837), : 53-124 (1838), : 125-166 (1839)
Fruhstorfer, 1908
Neue Argynnis und Melitaea Int. ent. Zs. 1 (41) : 310 (11 January)
Fruhstorfer, 1910
Neue palaearktische Rhopaloceren Ent. Zs. 24 (26) : 144 (1 October)
Fruhstorfer, 1917
Neue Rhopaloceren aus der Sammlung Leonhard Archiv Naturg. (A) 82 (2) : 1-28, pl. 1 (January)
Grose-Smith, 1887
Descriptions of two new Species of Butterflies from South Afghanistan Ann. Mag. nat. Hist. (5) 20 : 129-130
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
Grum-Grshimailo, 1893
Lepidoptera Palearctica Nova Horae Soc. ent. Ross. 27 (1-2): 127-129, : 379-386
Hemming, 1932
The butterflies of Transjordan Trans. ent. Soc. Lond. 80 : 269-299
Hemming, 1934
Some notes in the nomenclature of Palaearctic and African Rhopalocera Stylops 3 : 97-99
Hemming, 1941
On eight subspecies in the Lepidoptera Rhopalocera at present without valid names Proc. R. ent. Soc. Lond. 10 : 207-208
Higgins, 1938
A new Melitaeid butterfly from Asia Entomologist 71: 34
Higgins, 1941
An illustrated catalogue of the plaearctic Melitaea Trans. ent. Soc. Lond. 91 (7): 175-365, 16pls., 215figs
Higgins, 1955
A descriptive catalogue of Genus Mellicta Billberg and its species, with supplementary notes on the Genera Melitaea and Euphydryas Trans. R. ent. Soc. Lond. 106 (1): 1, 1-131, pl. 1-2, f. 1-66
Higgins, 1981
A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae Bull. Br. Mus. nat. Hist. (Ent.) 43 (3) : 77-243, f. 1-490,124a,125a
Huang, 2001
Report of H. Huang's 2000 Expedition to SE. Tibet for Rhopalocera Neue Entomologische Nachrichten, 51: 65-152
Huang & Murayama, 1992
Butterflies of Xinjiang province, China Tyō to Ga 43 (1) : 1-22
Hübner, [1826]; Hübner, [1799-1800]
Sammlung europäischer Schmetterlinge. I. Papiliones - Falter ("Erste Band") Samml. eur. Schmett. [1] : 3-74, : pl. 1-88, f. 1-446 (1799-1800), : pl. 89-96, f. 447-490 (1800-1803), : pl. 97-114, f. 491-588 (1803-1804), : pl. 115-119, f. 589-612 (1805), : pl. 120-124, f. 613-636 (1805-1806), : pl. 125-128, f. 637-652 (1806-1808), : pl. 129-144, f. 653-727 (1808-1813), : pl. 145-147, f. 728-741 (1814-1816), : pl. 148-150, f. 742-756 (1816-1817), : pl. 151, f. 757-762 (1818-1819), : pl. 152-154, f. 757-774 (1819-1822), : pl. 155, f. 775-778 (1822-1823), : pl. 156-161, f. 779-806 (1823), : pl. 162-175, f. 807-874 (1823-1824), : pl. 176, f. 869-874 (1824-1825), : pl. 177-181, f. 875-895 (1825-1826), : pl. 182-187, f. 896-981 (1827-1828), : pl. 188-195, f. 922-968 (1828-1832), : pl. 196, f. 969-974 (1832-1833), : pl. 197-201, f. 975-997 (1834-1836), : pl. 202-203, f. 998-1009 (1836-1838), : pl. 204-207, f. 1010-1029 (1837-1838)
Hübner, [1819]
Verzeichniss bekannter Schmettlinge, 1816-[1826] Verz. bek. Schmett. (1): [1-3], 4-16 (1816), (2): 17-32 (1819), (3): 33-48 (1819), (4): 49-64 (1819), (5): 65-80 (1819), (6): 81-96 (1819), (7): 97-112 (1819), (8): 113-128 (1819), (9): 129-144 (1819), (10): 145-160 (1819), (11): 161-176 (1819), (12): 177-192 (1820), (13): 193-208 (1820), (14): 209-224 (1821), (15): 225-240 (1821), (16): 241-256 (1821), (17): 257-272 (1823), (18): 273-288 (1823), (19): 289-304 (1823), (20): 305-320 (1825), (21): 321-336 (1825), (22): 337-352 (1825), (23-27): 353-431 ([1825])
Kolenati, 1846
Lepidoptera. In: Meletemata entomologica V. Insecta Caucasi Meletamata Entomologica 5 : 1-169, pl. 17-19
Kolesnichenko, 1999
A rewiew of the East-Palearctic tax of the Melitaea didyma (Esper, 1779)-group, 1. The M. ala Staudinger, 1881 - M. chitralensis Moore, 1901 complex Atalanta 30 (1/4): 87-117
Korshunov, 1972
Catalogue of the Butterflies (Lepidoptera, Rhopalocera) of USSR-fauna Ent. Obozr. 51 (2): 352-368
Krulikovsky, 1890
[russian: ? "experimental catalog of lepidoptera of Kazakh province"] Bull. Soc. imp. Nat. Moscou (n.s.) 4 (2) : 200-251, pl. 8
Lang, 1789
Verzeichniss seiner Schmetterlinge, in dem Gegenden um Augburg gesammelt und nach dem Wiener systematischen Verzeichniss eingetheilt, mit den Linneischen, auch deutschen und französischen Namen, und Anführung derjenigen Werke, worinn sie mit Farben abgebildet find Verz. Schmett. Gegend. Augsburg (ed. 2) : 1-226
Lederer, 1861
Über Alb. Kindermanns letzte lepidopterologische Ausbeute Wien. ent. Monats. 5 : 144-155
Leech, [1892]
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Lukhtanov, 1999
Neue Taxa und Synonyma zentalasiatischer Tagfalter Atalanta 30 (1/4): 135-150
Lukhtanov & Lukhtanov, 1994
Die Tagfalter Nordwestasiens Herbipoliana, 3: 440pp, 56pls, 400 figs
Matsumura, 1927
A list of the butterflies of Corea, with description of new species, subspecies and aberrations Insecta Matsumurana 1 (4) : 159-170
Ménétriés, 1859
Lépidoptčres de la Sibérie orientale et en particulier des rives de l'Amour Bull. phys-math. Acad. Sci. St. Pétersb. 17 (14) : 212-221, Mélanges biol. St.-Pétersb. 3 (1): 99-113 (1858)
Moore, 1865
List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2) : 486-509, pl. 30-31
Moore, 1874
List of Diurnal Lepidoptera collected in Cashmere Territory by Capt. R. B. Reed, 12th Regt., with Descriptions of new Species Proc. zool. Soc. Lond. 1874 (1) : 264-274, pl. 43
Murayama, 1955
New or Little Known Rhopalocera from China and Korea (with 10 text figures) Tyō to Ga 6 (1) : 1-4
Oberthür, 1876
Sur la Faune des Lépidoptčres de l'Algerie Étud. d'Ent. 1 : 1-74, pl. 1-4
Oberthür, 1909
Notes pour servir ą établir la Faune Francaise et Algeriénne des Lépidoptčres Étud. Lépid. Comp. 3 : 101-400, 405-415, pl. 11,16-20,23-26
Oberthür, 1915
Faune des Lépidoptčres de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315
Rothschild, 1913
[Expedition to the Central Western Sahara]. Lepidoptera Novit. Zool. 20 (1) : 109-143
Rothschild, 1917
Supplemental notes to Mr. Charles Oberthür's Faune des Lépidoptčres de la Barbarie, with lists of the specimens contained in the Tring Museum Novit. Zool. 24 (1): 61-120, : 325-373, : 393-409
Seitz, 1907
Die Großschmetterlinge des Palaearktischen Faunengebietes. 1. Die Palaearktischen Tagfalter Gross-Schmett. Erde 1 : 1-379
Shchetkin, 1981
The Nymphalidae of Surkhob valley from Tajikistan Ent. Obozr. Tajikistana: 159-173
Sheljuzhko, 1912
Eine neue Form von Melitaea didyma, O. Dt. ent. Z. Iris 26 (3) : 137-138
Sheljuzhko, 1914
Neue Rhopaloceren vom Pamir Dt. Ent. Z. Iris 28 (1) : 18-22
Sheljuzhko, 1929
Einige neue palaearktische Lepidopteren-Formen Mitt. Münch. Ent. Ges. 19 : 347-362, pl. 27-28
Sheljuzhko, 1929
Einige Bemerkunge über Melitaea ala Stgr. Mitt. Münch. Ent. Ges. 19 : 363-369, pl. 27-28
Sheljuzhko, 1935
Einige neue und wenig bekante Lepidopteren aus dem Westlichen Tian-Schan Mitt. Münch. ent. Ges. 25 : 27-38, pl. 3
Sodoffsky, 1837
Entomologische untersuchungen ueber die Gattungsnamen der Schmetterlinge Bull. Soc. imp. Nat. Moscou 1837 (6) : 76-97
Staudinger, 1861
Catalog der Lepidopteren Europas und der angrendzenden Länder in Staudinger & Wocke, Cat. Lep. (ed. 1): 1-192
Staudinger, 1870
Beitrag zur Lepidopterenfauna Griechenlands Horae Soc. ent. ross. 7 (1870) : 3-304, pl. 1-3
Staudinger, 1881
Beitrag zur Lepidopterenfauna Central-Asiens Stettin ent. Ztg 42 (7-9): 253-300, (10-12): 393-424
Staudinger, 1886
Centralasiatische Lepidopteren Stett. Ent. Ztg. 47 (4-6): 193-215, (7-9): 225-256
Staudinger, 1887
Centralasiatische Lepidopteren Stettin Ent. Ztg 48 (1-3) : 49-102
Staudinger, 1892
Die Macrolepidopteren des Amurgebiets. I. Theil. Rhopalocera, Sphinges, Bombyces, Noctuae in Romanoff, Mém. Lépid. 6 : 83-658, pl. 4-14
Staudinger, 1895
Ueber Lepidopteren von Uliassutai Dt. Ent. Z. Iris 8 (2) : 344-365 (1896)
Staudinger in Staudinger & Bang-Haas, 1882
Ueber einige neue Parnassius- und andere Tagfalter-Arten Central-Asiens Berl. ent. Z. 26 (1) : 161-177, pl. 1-2
Sushkin, 1913
Zur anatomischen Begründung einiger palaerktischer Arter der Gattung Melitaea Z. wiss. Insektenbiologie 9 (6/7) : 169-175
Turati, 1919
Nuove forme di lepidotteri Naturalista sicil. 23 (7-12) : 203-351, pl. 1-4
Tytler, 1926
Notes on some new and interesting Butterflies from India and Burma. (1) & (2) J. Bombay nat. Hist. Soc. 31 : 248-260, (3): 578-590, pl. 1-5 [?]
Verity, 1919
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 31 : 26-31,43-48,87-89,121-129,178-184,193-201
Verity, 1929
The Asiatic origins of the Westenr Palearctic Rhopalocera exemplified by Melitaea didyma Esp. Ent. Rec. J. Var. 41 : 31-34, 39-43, 63-67, 72-76, 89-91, 111-118, 127-132
Wagner, 1913
Beitrag zur Lepidopterenfauna des Iligebietes sowie des Sary-Dschas (Asia centr.) Ent. Mitt. 2 (1): 22-30, (2): 50-62, (3): 88-95, (4): 112-126, (5): 153-158, (6): 185-190, (7/8): 244-254, (9): 285-288
Weiss & Major, 2000
A new species of the genus Melitaea Fabricius, 1807 from Iran Atalanta 31 (1/2): 123-128
Wiltshire, 1941
New lepidoptera from S.W.Iran J. Bomb. Nat. Hist. Soc. 42 (3): 472-477
Wyatt, 1961
Additions to the Rhopalocera of Afghanistan with descriptions of new species and subspecies J. Lep. Soc. 15 (1): 1-18, 3pls


If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.