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Hadula Staudinger, 1889
Hadula Staudinger, 1889; Stettin Ent. Ztg 50 (1-3) : 43, TS: Hadula insolita Staudinger
= Anarta ; [NE9, 233]
Discestra Hampson, 190515; Mamestra chartaria, Grote
= ; [NE4, 21]
= Anarta ; [NE9, 233]
Aglossestra Hampson, 1905; 225, TS: Aglossestra deserticola Hampson
= ; [NE4, 21]
= Anarta ; [NE9, 233]
Salacia Boie, 1839; 391 (preocc. Salacia Lamouroux, 1816 (Coelenterata)), TS: Mamestra sodae Boisduval
= Discestra ; [NACL, 148]
= Discertra ; [POOLE]
= ; [NE4, 21]
Cardiestra Boursin, 1963; 86, TS: Mamestra eremistis Püngeler
= ; [NE4, 21]
Melanarta Beck, 1991; Atalanta 22 (2/4): 189, TS: Noctua melanopa Thunberg
= ; [NE4, 21]
Calocestra Beck, 1991; Atalanta 22 (2/4): 190, TS: Noctua pugnax Hübner
= ; [NE4, 21]
Trifcestra Beck, 2000
= ; [NE4, 21]
Dianthcestra Beck, 2000
= ; [NE4, 21]

Dicestra ; McDunnough, 1937, (missp.)
Discestra (Hadenini) ; [NACL, 148]
Aglossestra (Hadeninae) ; [POOLE]
 Cardiestra (Hadeninae) ; [POOLE]
Hadula (Hadeninae) ; [POOLE]
Hadula (Hadenini) ; [FE]
Hadula (Discestrina) ; [NE4, 18, 21]

Subgenus Hadula Staudinger, 1889
Hadula insolita Staudinger, 1889
Turkestan - Mongolia. See [About maps] Hadula insolita Staudinger, 1889; 43, TL: South of Issyk-kul
Hadula insolita ; [POOLE]
Hadula (Hadula) insolita ; [NE4, 21]
Anarta (Hadula) insolita ; [NE12, 184, pl. 14, f. 11-15]
H. i. uigurica Hacker, 1998
Hadula (Hadula) insolita uigurica ; [NE4, 21]
Hadula anthracina (Hacker, 1996)
Hadula (Hadula) anthracina ; [NE4, 21]
Hadula gandhara Hacker, 1998
Hadula (Hadula) gandhara ; [NE4, 21]
Hadula nekrasovi Gyulai & Varga, 1998
Hadula (Hadula) nekrasovi ; [NE4, 21]
Hadula insperata Gyulai & Hacker, 1998
Hadula (Hadula) insperata ; [NE4, 21]
Hadula ultramontana Gyulai & Hacker, 1998
Hadula (Hadula) ultramontana ; [NE4, 21]
Hadula sabulorum (Alphéraky, 1882)
-. See [About maps] Mamestra sabulorum Alphéraky, 1882; 69, pl. 3, f. 58, TL: Kouldja district, Korgosse
Hadula sabulorum ; [POOLE]
H. s. distincta (Staudinger, 1889)
Mamestra sabulorum var. distincta Staudinger, 1889; 35, TL: South of Ikkyk-Kul
= Hadula sabulorum ; [POOLE]
Hadula leucheima Boursin, 1963; 43, pl. 11, f. 1-2, TL: Issyk-kul
Hadula leucheima ; [POOLE]
Hadula (Hadula) sabulorum distincta ; [NE4, 21]
H. s. rhodina (Christoph, 1893)
Mamestra sabulorum var. rhodina Christoph, 1893; 91, TL: Ordubad
= Hadula sabulorum ; [POOLE]
Hadula (Hadula) sabulorum rhodina ; [NE4, 21]
H. s. segnis (Püngeler, 1906)
Xinjiang, ... Scotogramma (Mamestra) segnis Püngeler, 1906; 90, pl. 6, f. 17, TL: [Xingjiang] Lob-Noor
Hadula segnis ; [POOLE]
Hadula (Hadula) sabulorum segnis ; [NE4, 21]
H. s. pulverata (Bang-Haas, 1907)
Malta Mamestra pulverata Bang-Haas, 1907; 71, pl. 3, f. 8, TL: Tunisia, Gafsa
Polia cinnamomeogrisea Rotschild, 1913; 121, TL: "Central Western Sahara, Bordj Chegga"
= Hadula pulverata ; [POOLE]
chiamaera (Rothschild, 1920)
Hadula pulverata ; [POOLE]
Hadula (Hadula) sabulorum pulverata ; [NE4, 18, 21]
Hadula engedina Hacker, 1998
Hadula (Hadula) engedina ; [NE4, 21]
Hadula halodeserti Varga, 1973
Mongolia. See [About maps] Hadula halodeserti Varga, 1973; 212, pl. 7, f. 2, TL: Mongolia, South Gobi Aimak, Ablussloses Becker
Hadula halodeserti ; [POOLE]
Hadula (Hadula) halodeserti ; [NE4, 21]

Subgenus ?
Hadula brassicina (Draudt, 1934)
Altai, .... See [About maps] Scotogramma brassicina Draudt, 1934; 98, pl. 14, row c, TL: Western Altai
Hadula brassicina ; [POOLE]
Hadula chiklika (Moore, 1878)
Taeniocampa chiklika Moore, 1878; 234, TL: Yarkund, SE of Chiklik
Hadula chiklika ; [POOLE]
Hadula chimaera (Rothschild, 1920)
Algeria, .... See [About maps] Scotogramma chimaera Rothschild, 1920; 57, pl. 17, f. 6, TL: Algeria, Ain Sefra
Hadula chimaera ; [POOLE]
Hadula gigantea (Rebel, 1909)
Libya, Algeria, Egypt. See [About maps] Thargelia gigantea Rebel, 1909; 132, TL: Sinai, Ajun Mua Oasis; W. Werdan
Odontelia griseola Rothschild, 1913; Novit. Zool. 20 (1) : 121, TL: [Algeria] Mrraier, halfway Biskra and Touggourt; halfway Ouargla and El-Golea
= ; [POOLE]
Odontelia grazianii Krüger, 1933; 95, f. 2, TL: Cirenaica, Maaten, Giofer
= ; [POOLE]
Hadula gigantea ; [POOLE]
Hadula ochrea (Warren, 1909)
Xinjiang, .... See [About maps] Odontelia ochrea Warren, 1909; 81, pl. 20, row b, TL: [Xinjiang] Kuldja
Hadula ochrea ; [POOLE]
Hadula orbona Bang-Haas, 1912
Hadula orbona Bang-Haas, 1912; Dt. ent. Z. Iris 26 (3): 151, (4): pl. 6, f. 15, TL: Kragai-tau
Hadula orbona ; [POOLE]

Subgenus Cardiestra Boursin, 1963
Hadula eremistis (Püngeler, 1904)
Mamestra eremistis Püngeler, 1904; 130, TL: Merw
Cardiestra eremistis ; [POOLE]
Hadula (Cardiestra) eremistis ; [NE4, 21]
H. e. avicenna hacker, 1998
Hadula (Cardiestra) eremistis avicenna ; [NE4, 21]
Hadula vaciva (Püngeler, 1906)
Dicestra[sic] vaciva Püngeler, 1906; 90, pl. 6, f. 4, TL: Ili region
Cardiestra vaciva ; [POOLE]
Hadula (Cardiestra) vaciva ; [NE4, 21]
Hadula gobideserti (Varga, 1973)
Mongolia. See [About maps] Cardiestra gobideserti Varga, 1973; 209, pl. 7, f. 1, TL: Mongolia, South Gobi Aimak, Ablussloses Becken, 10km NNE of Dalanzargad
Cardiestra gobideserti ; [POOLE]
Hadula (Cardiestra) gobideserti ; [NE4, 21]
Hadula halolimna (Gyulai & Varga, 1998)
Hadula (Cardiestra) halolimna ; [NE4, 18, 21]
Hadula parvula Hacker, 1998
Hadula (Cardiestra) parvula ; [NE4, 21]
Hadula vassilinini (O. Bang-Haas, 1927)
Transcaucasus, .... See [About maps] Dicestra[sic] vassilinini O. Bang-Haas, 1927; 85, pl. 10, f. 25, TL: Transcuacasus, Elisabethpol
Cardiestra vassilini ; [POOLE]
Hadula (Cardiestra) vassilinini ; [NE4, 21]

Subgenus Ptochicestra Hacker, 1998
Hadula ptochica (Püngeler, 1900)
Mamestra ptochica Püngeler, 1900; 104, TL: "Central Asia"
Hadula ptochica ; [POOLE]
Hadula (Ptochicestra) ptochica ; [NE4, 21]
H. p. transcaucasica Hacker, 1998
Hadula (Ptochicestra) ptochica transcaucasica ; [NE4, 21]
Hadula osmana Hacker, 1998
Hadula (Ptochicestra) osmana ; [NE4, 21]

Subgenus Aglossestra Hampson, 1905
Hadula deserticola (Hampson, 1905)
coastal (Tunisia, Algeria, N.Egypt). See [About maps] Aglosesstra deserticola Hampson, 1905; 225, f. 38, TL: Egypt, Suez
Hadula mariaeludovicae Lucas, 1914; 311, TL: Tunis
= ; [POOLE]
Derthisa affinis Rothschild, 1914; 328, TL: Algeria, Guelt-es-Stel
= ; [POOLE]
Discestra deserticola ;
Aglossestra deserticola ; [POOLE]
Hadula (Aglossestra) deserticola ; [NE4, 21]
Anarta (Aglossestra) deserticola ; [NE9, 233, pl. 11, f. 46-50, (gen.) f. 142, 284]

Subgenus Calocestra Beck, 1991
Hadula pugnax (Hübner, [1824])
SEU. See [About maps] Noctua pugnax Hübner, [1824]; pl. 156, f. 726-727, TL: Europe
Noctua treitschkii Boisduval, 1827; Annls Soc. linn. Paris 6: 111, TL: Languedoc, [S.France]
= ; [POOLE]
Scotogramma treitschkii f. petricolor Turati, 1923; 53, TL: Modena
= ; [POOLE]
Scotogramma treitschkii italica Berio, 1942; 125, TL: Italy, Savona Province, Cengio
= ; [POOLE]
Discestra pugnax, Moths and Butterflies of Europe [MBE]
 Hadena treitschkei[sic?] ; (missp.?)
Scotogramma treischkei[sic] ; Rothschild, 1915, Novit. Zool. 22 (2): 230
Discestra pugnax ; [POOLE]
Hadula (Calocestra) pugnax ; [NE4, 18, 21]

Larva on Hippocrepis, Lotus Forster & Wohlfahrt, 1971, [POOLE]

H. p. intermedia (Pinker, 1980)
Asia Minor Discestra pugnax intermedia Pinker, 1980; 70, pl. 1, f. C1, TL: Asia Minor, Gürün
= Discestra pugnax ; [POOLE]
Hadula (Calocestra) pugnax intermedia ; [NE4, 21]
Hadula atlantis (Schwingenschuss, 1955)
Morocco, .... See [About maps] Scotogramma pugnax atlantis Schwingenschuss, 1955; 227, TL: Morocco, Atlas Mntns
= Discestra pugnax ; [POOLE]
Hadula (Calocestra) atlantis ; [NE4, 21]
Hadula latemarginata (Wiltshire, 1976)
Discestra trifolii latemarginata Wiltshire, 1976; 75, pl. 1, f. 5a-b, TL: Turkey, Pontic Mts., Zigana Pass
= Discestra trifolii ; [POOLE]
Hadula (Calocestra) latemarginata ; [NE4, 21]
Hadula baksana Schintlmeister & Poltawski, 1986
Hadula (Calocestra) baksana ; [NE4, 21]
Hadula odontites (Boisduval, 1829)
SEU, CEU. See [About maps] Hadena odontites Boisduval, 1829; Eur. Lepid. Index Meth.: 71, TL: Switzerland
= Hada nana ; [POOLE]
nana; , TL: Hufnagel, 1766##
microdon; , TL: Guenée, 1852##
Mamestra maromorsa var. dalmatina Schwingenschuss, 1926; 10, TL: Dalmatia
= Discestra microdon ; [POOLE]
Hadula (Calocestra) odontites ; [NE4, 18, 21]
Misidentified/misapplied:
marmorosa; Borkhausen, 1972
?Discestra microdon; [POOLE]
H. o. boisduvali Hacker, 1998
Hadula (Calocestra) odontites boisduvali ; [NE4, 21]
H. o. turkestana Hacker, 1998
?rosea Guylai & Ronkay, 1998
Hadula (Calocestra) odontites turkestana ; [NE4, 21]
Hadula schawyra (Bang-Haas, 1927)
Scotogramma schawyra Bang-Haas, 1927; 85, pl. 10, f. 26, TL: Tannuola Mtns, Schawyra
Discestra schawyra ; [POOLE]
Hadula (Calocestra) schawyra ; [NE4, 21]
Hadula schneideri (Staudinger, 1900)
Xinjiang, .... See [About maps] Mamestra schneideri Staudinger, 1900; 368, TL: [Xinjiang] Korla
Discestra schneideri ; [POOLE]
Hadula (Calocestra) schneideri ; [NE4, 21]
H. s. herczigi Hacker, 1998
Hadula (Calocestra) schneideri herczigi ; [NE4, 22]
Hadula eversmanni (Staudinger, 1900)
Hadula (Calocestra) eversmanni ; [NE4, 22]
Hadula furcula (Staudinger, 1889)
Mamestra furcula Staudinger, 1889; 36, TL: Issyk-kul
Discestra furcula ; [POOLE]
Hadula (Calocestra) furcula ; [NE4, 22]
H. f. tibetica Hacker, 1998
Hadula (Calocestra) furcula tibetica ; [NE4, 22]
Hadula perdentata (Hampson, 1894)
Kashmir, .... See [About maps] Hadena perdentata Hampson, 1894; 198, TL: Kashmir
Discestra perdentata ; [POOLE]
Hadula (Calocestra) perdentata ; [NE4, 22]
H. p. bifida (Püngeler, 1902)
Mamestra bifida Püngeler, 1902; 149, pl. 5, f. 3, TL: Alexander Mnts
= Discestra perdentata ; [POOLE]
Hadula (Calocestra) perdentata bifida ; [NE4, 22]
Hadula vargai (Hreblay & Ronkay, 1998)
Hadula (Calocestra) vargai ; [NE4, 22]
H. v. madefacta Hacker, 1998
Hadula (Calocestra) vargai madefacta ; [NE4, 22]
Hadula furca (Eversmann, 1852)
Hadena furca Eversmann, 1852; 154, TL: Eastern Siberia
Discestra furca ; [POOLE]
Hadula (Calocestra) furca ; [NE4, 22]
Hadula colletti (Sparre-Schneider, 1876)
Norway, ... See [About maps] Dianthoecia colletti Sparre-Schneider, 1876; 56, TL: Norway, Dovre Alps, Domaas
= Discestra furca ; [POOLE]
Hadula (Calocestra) colletti ; [NE4, 18, 22]
H. c. originalis Hacker, 1998
Hadula (Calocestra) colletti originalis ; [NE4, 22]
Hadula farnhami (Grote, 1873)
Mamestra farnhami Grote, 1873; 103, pl. 3, f. 2, TL: Colorado Territory
Discestra farnhami, Butterflies and Moths of North America [BAMONA]
Discestra farnhami, Noctuid Search (record) [NOCTS]
 Discestra farnhami ; [POOLE]; [NACL, #10232]
Hadula (Calocestra) farnhami ; [NE4, 22]
H. f. palaearctica Hacker, 1998
Hadula (Calocestra) farnhami palaearctica ; [NE4, 18, 22]
H. f. shandura Hacker, 1998
Hadula (Calocestra) farnhami shandura ; [NE4, 22]
Hadula imperspicua Hacker, 1998
Hadula (Calocestra) imperspicua ; [NE4, 18,22]
H. i. mandschurica Hacker, 1998
Hadula (Calocestra) imperspicua mandschurica ; [NE4, 22]
Hadula hampsoni Hacker, 1998
Hadula (Calocestra) hampsoni ; [NE4, 22]
Hadula mendica (Staudinger, 1895)
Turkey, .... See [About maps] Mamestra mendica Staudinger, 1895; 272, pl. 9, f. 13, TL: [Turkey] Hadjin
Discestra mendica ; [POOLE]
Hadula (Calocestra) mendica ; [NE4, 22]
H. m. mendosa Hacker, 1998
Hadula (Calocestra) mendica mendosa ; [NE4, 22]
Hadula mendax (Staudinger, 1879)
SEEU. See [About maps] Hadula (Calocestra) mendax ; [NE4, 22]
H. m. occidentalis Hacker, 1998
Hadula (Calocestra) mendax occidentalis ; [NE4, 18,22]
H. m. phoenicica Hacker, 1998
Hadula (Calocestra) mendax phoenicica ; [NE4, 22]
Hadula hoplites (Staudinger, 1901)
Mamestra armata var. hoplites Staudinger, 1901; 160, TL: Tura
Discestra loeffleri Reisser, 1958; 553, pl. 5, f. 8, pl. 6, f. 3, TL: Iran, between Isfahan and Gom
Discestra loeffleri ; [POOLE]
Discestra hoplites ; [POOLE]
Hadula (Calocestra) hoplites ; [NE4, 18, 22]
Hadula armata (Staudinger, 1888)
Mamestra armata Staudinger, 1888; 15, TL: Samarkand
Discestra armata ; [POOLE]
Hadula (Calocestra) armata ; [NE4, 22]
H. a. balkhashensis Hacker, 1998
Hadula (Calocestra) armata balkhashensis ; [NE4, 22]
Hadula isoloma (Püngeler, 1904)
Mamestra isoloma Püngeler, 1904; 286, pl. 6, f. 2, TL: "Sarafschan region"
Discestra isoloma ; [POOLE]
Hadula (Calocestra) isoloma ; [NE4, 22]
Hadula actinobola (Eversmann, 1837)
Hadena actinobola Eversmann, 1837; Bull. Soc. imp. Nat. Moscou 1837 (6) : 38, TL: Caucasus
= Discestra dianthi ; [POOLE]
Hadula nupponenorum Hacker & Fibiger, 2002
= Anarta actinibola ; [NE12, 185]
Hadula (Calocestra) nupponenorum ; [NE4, 18, 22]
Anarta (Hadula) actinibola ; [NE12, 185, pl. 14, f. 6-7]
Hadula dianthi (Tauscher, 1809)
SEU, S.Urals, Caucasus, .... See [About maps] Noctua dianthi Tauscher, 1809; 316, pl. 20, f. 2, TL: Russia, Sarepta; Waskuntschatkoi
Hadena dentigera Eversmann, 1837; 35, TL: Urals; Saratschik
= ; [POOLE]
furculoides F. Wagner, 1930
Discestra dianthi ; [POOLE]
Hadula (Calocestra) dianthi ; [NE4, 22]

Larva on Aster trifolium Forster & Wohlfahrt, 1971, [POOLE]

H. d. dianthi
Hadula (Calocestra) dianthi dianthi ; [NE4, 18, 22]
H. d. lukhtanovorum Hacker, 1998
Hadula (Calocestra) dianthi lukhtanovorum ; [NE4, 22]
H. d. hungarica (F. Wagner, 1930)
Hadula (Calocestra) dianthi hungarica ; [NE4, 18, 22]
H. d. parenzani Hacker, 1998
Hadula (Calocestra) dianthi parenzani ; [NE4, 18, 22]
H. d. hubiesi (Lajonquiére, 1969)
Spain, ... Discestra dianthi hubiesi Lajonquiére, 1969; 158, pl. 11, f. 4, TL: Spain, Zaragoza, Bujaraloz
= Discestra dianthi ; [POOLE]
Hadula (Calocestra) dianthi hubiesi ; [NE4, 18, 22]
H. d. getula (Rungs, 1942)
Morocco, ... Scotogramma dianthi getula Rungs, 1942; 162, pl. 1, f. 12, TL: Morocco, Moyen Valley of Dra, Mader Bergat
= Discestra dianthi ; [POOLE]
Hadula (Calocestra) dianthi getula ; [NE4, 22]
fiHadula trifolii (Hufnagel, 1766)
EU, Niger, Tunisia, Iran, Quebec, .... See [About maps] Savikkayökkönen Nutmeg Clover Cutworm Meldenflureule Kleefeldeule
Phalaena trifolii Hufnagel, 1766; 398, TL: Berlin region
Noctua chenopodii Denis & Schiffermüller, 1775; Ankündung eines systematischen Werkes von den Schmetterlingen der Wienergegend: 82, TL: Vienna region
= ; [POOLE]
Phalaena (Noctua) verna Esper, 1787; Die Schmett., Th. IV, Bd. 1 (42-44): 272, (35): pl. 117, f. 5-6 (Noct. 38a), TL: Leipzig region
= ; [POOLE]
Phalaena (Noctua) saucia Esper, 1790; Die Schmett., Th. IV, Bd. 1 (41): pl. 152, f. 5 (Noct. 73), TL: Tyrol
= ; [POOLE]
Phalaena (Noctua) saucia Esper, 1804; Die Schmett., Th. IV, Bd. 2 (Abs. 1) (52): 505
Noctua infraina Haworth, 1809; 192, TL: Great Britain
= ; [NACL, #10223]; [POOLE]
Noctua contribulis Duponchel, 1827; 357, pl. 122, f. 1, TL: Montpellier
= ; [POOLE]
Orthosia farkasii Treitschke, 1835; Schmett. Eur. 10 (2): 74, TL: "Shrmien"
= ; [POOLE]
Apamea inquieta Walker, 1857; 730
= ; [NACL, #10223]; [POOLE]
Hadena intermissa Walker, 1857; 587, TL: Chile
= ; [POOLE]
Apamea glaucovaria Walker, 1860; 255, TL: Quebec, Montreal
= ; [NACL, #10223]; [POOLE]
Mamestra trifolii var. major Speyer, 1875; 138, TL: United States
= ; [POOLE]; [NACL, #10223]
Mamestra canescens Moore, 1878; 233, TL: [Tibet] Yarkund, Kargalik
= ; [POOLE]
Scotogramma cinnamomina Rothschild, 1913; 121, TL: Central Western Sahara, Nza-Ben-Rzi [Niger]
= ; [POOLE]
Cardepia taylori Rothschild, 1921; 171, TL: Mesopotamia, Amara
= ; [POOLE]
Scotogramma trifolii var. fructicosae Dumont, 1925; 334, TL: Tunisia, Tozeur
= ; [POOLE]
Scotogramma trifolii f. zermattensis Draudt, 1934; 96, TL: Zermatt
= ; [POOLE]
chenopodii Denis & Schiffermüller, 1775
Discestra trifolii, Butterflies and Moths of North America [BAMONA]
Discestra trifolii, Lynn Scott's Lepidoptera Images [Lynn Scott]
Hadula trifolii, Svenska fjärilar [SVF]
Discestra trifolii, Noctuid Search (record) [NOCTS]
Discestra trifolii, Moths and Butterflies of Europe [MBE]
Discestra trifolii, UK Moths [Ian Kimber]
 Discestra trifolii ; [NACL, #10223]; [POOLE]
Hadula (Calocestra) trifolii ; [NE4, 18, 22]
[trifolii-1.jpg] 623x482 (~68Kb) Germany, Baden-Württemberg, Schmie, Schafäcker (8°49'E, 48°59'N, 300m), 04.06.2000, Photo © Karl Hofsäß [trifolii-2.jpg] 943x703 (~115Kb) Russia, Moscow area, 18.8.2008, Photo © D. Smirnov [trifolii.jpg] 500x533 (~39Kb) FINLAND: Ka: Virolahti, 671:53, m 29-4.8.1978, f 6-10.8.1972, Markku Savela leg.

Flight from June to first half of July. Second generation from latter half of July to September.

The second generation is much more abundant than the first.

Larva on Allium cepa, Chenopodium spp., C. album, Atriplex spp.


H. t. albifusa (Walker, 1857)
Nova Scotia, ... Hadena albifusa Walker, 1857; 753, TL: Nova Scotia
= Discestra trifolii ; [POOLE]
Discestra trifolii albifusa ; [NACL, #10223a]
Hadula gredosi (Laever, 1977)
Spain. See [About maps] Discestra gredosi Laever, 1977; 74, f. 5, TL: Spain, Gredos
= Discestra trifolii ; [POOLE]
Hadula (Calocestra) gredosi ; [NE4, 18, 22]
Anarta (Calocestra) gredosi ; [NE12, 184, pl. 14, f. 8-10]
Hadula stigmosa (Christoph, 1887)
SEU. See [About maps] Mamestra stigmosa Christoph, 1887; 70, pl. 3, f. 12, TL: Salz Steppes
Discestra stigmosa ; [POOLE]
Hadula (Calocestra) stigmosa ; [NE4, 22]
H. s. stigmosa
Hadula (Calocestra) stigmosa stigmosa ; [NE4, 18]
H. s. atlantica (Boursin, 1936)
France, Corsica, Italy, ... Scotogramma stigmosa atlantica Boursin, 1936; 154, TL: France, La Faute
= Discestra stigmosa ; [POOLE]
Discestra stigmosa corsicola Boursin, 1962; 302, TL: Corsica, St.-Florent
= Discestra stigmosa ; [POOLE]
Dicestra[sic] stigmosa parentii Berio, 1980; 225, TL: Apulia, Manfrednia, Ippocampo
= Discestra stigmosa ; [POOLE]
Dicestra[sic] stigmosa meridionalis Berio, 1980; 226, TL: Plugia, Mar Piccolo, Taranto
= Discestra stigmosa ; [POOLE]
Dicestra[sic] stigmosa vicarioi Berio, 1980; 226, TL: [Italy] Grosseto, Uccellina
= Discestra stigmosa ; [POOLE]
Hadula (Calocestra) stigmosa atlantica ; [NE4, 18, 22]
Hadula sodae (Boisduval, 1829)
SEU. See [About maps] Mamestra sodae Boisduval, 1829; (errata & addenda) 5, TL: Monspelii
Discestra sodae ; [POOLE]
Hadula (Calocestra) sodae ; [NE4, 18, 22]
H. s. rosacea (Rothschild, 1920)
Algeria, ... Scotogramma sodae rosacea Rothschild, 1920; 56, TL: Algeria, Biskra
= Discestra sodae ; [POOLE]
sodae var. maroccana Rungs, 1938
Hadula (Calocestra) sodae rosacea ; [NE4, 22]
H. s. salicorniae (Dumont, 1925)
Tunisia, ... Scotogramma salicorniae Dumont, 1925; 332, TL: Tunisia, Tozeur
= Discestra sodae ; [POOLE]
Scotogramma salicorniae var. raselaini Dumont, 1925; 333, TL: Tunisia, Ras el Aioun Rozeur
= Discestra sodae ; [POOLE]
Scotogramma picta Turati, 1926; 37, f. 4, TL: Cirenaica, Tobruk
= Discestra sodae ; [POOLE]
Hadula (Calocestra) sodae salicorniae ; [NE4, 22]
H. s. fuerteventuriensis (Pinker & Bacallado, 1979)
Canary Islands Discestra sodae fuerteventuriensis Pinker & Bacallado, 1979; 86, TL: Fuerteventura, Jandia Playa
= Discestra sodae ; [POOLE]
Hadula (Calocestra) sodae fuerteventuriensis ; [NE4, 22]
Hadula explicata (Sukhareva, 1976)
Hadula (Calocestra) explicata ; [NE4, 22]
fiHadula melanopa (Thunberg, 1791)
N.Scandinavia, Alps?. See [About maps] Paljakan kangasyökkönen Broad-bordered White Underwing
Noctua melanopa Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 42, TL: "Lapponia"
Noctua alpicola Quenzel, 1802; 253, pl. 1, f. 5-6, TL: "Lappland"
= ; [NACL, #10327]; [POOLE]
Anarta wistroemi Lampa, 1885
= ; [NACL, #10327]
Anarta melanopa, Butterflies and Moths of North America [BAMONA]
Hadula melanopa, Svenska fjärilar [SVF]
Discestra melanopa, Moths and Butterflies of Europe [MBE]
Anarta melanopa, UK Moths [Ian Kimber]
 Anarta melanopa ; [NACL, #10327]; [POOLE]
Hadula (Calocestra) melanopa ; [NE4, 22]
[melanopa-1f.jpg] 800x614 (~103Kb) female FINLAND: Li: Utsjoki Ailigas, 775:50, 5.7.1997, Photo © Markku Savela [melanopa.jpg] 500x490 (~36Kb) FINLAND: Li: Utsjoki, 775:50, m 12.7.1996, Le: Kilpisjärvi, 767:25, f 17.7.1982, Markku Savela leg.

Flight from first half of June to first half of July.

Larva on Polygonum viviparum, Empetrum, Vaccinium uliginosum, Dryas


H. m. melanopa
Hadula (Calocestra) melanopa melanopa ; [NE4, 18]
H. m. rupestralis (Hübner, [1799])
Pyralis rupestralis Hübner, [1799]; pl. 21, f. 138, TL: Europe
= Anarta melanopa ; [NACL, #10327]; [POOLE]
Noctua vidua Hübner, [1808]; pl. 86, f. 403, TL: Europe
= Anarta melanopa ; [NACL, #10327]; [POOLE]
Noctua tristis Hübner, [1809]; pl. 95, f. 446 (preocc. Noctua tristis Fabricius, 1775), TL: Europe
= Anarta melanopa ; [NACL, #10327]; [POOLE]
Noctua rupestris Hübner, [1819]; pl. 141, f. 644-645 (emend. Pyralis rupestralis Hübner, [1799]), TL: Europe
= Anarta melanopa ; [NACL, #10327]; [POOLE]
Hadula (Calocestra) melanopa rupestralis ; [NE4, 18, 22]
H. m. brunnea (Tutt, 1892)
Hadula (Calocestra) melanopa brunnea ; [NE4, 18]
H. m. koizumidakeana (Matsumura, 1927)
Japan Anarta koizumidakeana Matsumura, 1927; Ins. Matsumurana 1 (3) : 116, f. 6, TL: Hokkaido, Kurodake & Koizumidade
= Anarta melanopa ; [POOLE]
Hadula (Calocestra) melanopa koizumidakeana ; [NE4, 22]
Hadula nigrolunata (Packard, 1867)
Labrador, Yukon, British Columbia, New Hampshire. See [About maps] Anarta nigrolunata Packard, 1867; 40, TL: Labrador, Okak
= Anarta melanopa ; [POOLE]; [NACL, #10327]
Anarta nigrolunata, Moth Photographers Group [MPG]
Anarta nigrolunata, E.H. Strickland Ent. Mus, U of [ALBERTA]
 "Anarta" nigrolunata ; Lafontaine & Troubridge
?H. n. laerta (Smith, 1903)
Utah, ... Anarta melanopa laerta Smith, 1903
Anarta laerta Smith, 1903; J. N.Y. Ent. Soc. 11 (1) : 20, TL: Utah, Silver Lake
= Anarta melanopa ; [POOLE]
Anarta melanopa laerta ; [NACL, #10327a]
Hadula militzae (Kozhantschikov, 1947)
Alai, ... Anarta militzae Kozhantschikov, 1947; 16, pl. 1, f. 3, TL: South Alai, Dzhaidak
Anarta militzae ; [POOLE]
Hadula (Calocestra) militzae ; [NE4, 22]
Hadula inexpecta (Yoshimoto, 1995)
Hadula (Calocestra) inexpecta ; [NE4, 22]
Hadula hreblayi Hacker, 1998
Hadula (Calocestra) hreblayi ; [NE4, 22]

Subgenus Pulchrohadula Hacker, 1998
Hadula triphaenopsis (Oberthür, 1893)
Hadula (Pulchrohadula) triphaenopsis ; [NE4, 22]

Subgenus ?

The "NA" species-group

Were in Discestra, may belong else...

Hadula chartaria (Grote, 1873)
California, .... See [About maps] Mamestra chartaria Grote, 1873; 138, pl. 4, f. 12, TL: California
Discestra chartaria, Butterflies and Moths of North America [BAMONA]
Discestra chartaria, Noctuid Search (record) [NOCTS]
 Discestra chartaria ; [POOLE]; [NACL, #10222]
Hadula mutata (Dod, 1913)
Alberta, .... See [About maps] Mamestra mutata Dod, 1913; 29, TL: Alberta, Calgary
Discestra mutata, Butterflies and Moths of North America [BAMONA]
Discestra mutata, Noctuid Search (record) [NOCTS]
Discestra mutata, The Moths of Southeastern Arizona [Bruce Walsh]
 Discestra mutata ; [NACL, #10224]; [POOLE]
Hadula fulgora (Barnes & McDunnough, 1918)
California, .... See [About maps] Scotogramma fulgora Barnes & McDunnough, 1918; 94, pl. 16, f. 9, TL: California, Inyo Co., Olancha
Discestra fulgora, Butterflies and Moths of North America [BAMONA]
Discestra fulgora, Noctuid Search (record) [NOCTS]
 Discestra fulgora ; [NACL, #10225]; [POOLE]
Hadula castrae (Barnes & McDunnough, 1912)
New Mexico, ... See [About maps] Scotogramma castrae Barnes & McDunnough, 1912; 11, pl. 1, f. 11, TL: New Mexico, F. Wingate
Discestra castrae, Butterflies and Moths of North America [BAMONA]
 Discestra castrae ; [NACL, #10226]; [POOLE]
H. c. ultra (Barnes & Benjamin, 1924)
New Mexico, ... Scotogramma castrae f. ultra Barnes & Benjamin, 1924; 123, TL: New Mexico, Jemez Springs
= Discestra castrae ; [POOLE]
Discestra castrae ultra ; [NACL, #10226a]
Hadula hamata (McDunnough, 1930)
Montana, .... See [About maps] Scotogramma hamata McDunnough, 1930; 183, f. 3, TL: Montana, Hamilton
Discestra hamata, Butterflies and Moths of North America [BAMONA]
Discestra hamata, Noctuid Search (record) [NOCTS]
 Discestra hamata ; [NACL, #10227]; [POOLE]
Hadula oregonica (Grote, 1881)
Oregon, .... See [About maps] Mamestra trifolii var. oregonica Grote, 1881; 230, TL: Oregon
Discestra oregonica, Butterflies and Moths of North America [BAMONA]
Discestra oregonica, Noctuid Search (record) [NOCTS]
 Discestra oregonica ; [POOLE]; [NACL, #10228]
H. o. morana (Smith, 1910)
Wyoming, ... Mamestra morana Smith, 1910; 361, pl. 11, f. 6, TL: Wyoming, Yellowstone Park
= Discestra oregonica ; [POOLE]
Discestra oregonica morana ; [NACL, #10228a]
H. o. columbica (McDunnough, 1930)
British Columbia, ... Scotogramma oregonica var. columbica McDunnough, 1930; 181, f. 1, TL: British Columbia, Keremeos
= Discestra oregonica ; [POOLE]
Discestra oregonica columbica ; [NACL, #10228b]
H. o. montanica (McDunnough, 1930)
Montana, ... Scotogramma oregonica var. montanica McDunnough, 1930; 182, f. 2, TL: Montana, Bozeman
= Discestra oregonica ; [POOLE]
Discestra oregonica montanica ; [NACL, #10228c]
Hadula alta (Barnes & Benjamin, 1942)
Alberta, .... See [About maps] Scotogramma alta Barnes & Benjamin, 1942; 123, TL: Alberta, Nordegg
Discestra alta, Butterflies and Moths of North America [BAMONA]
 Discestra alta ; [NACL, #10229]; [POOLE]
Hadula subalbida (Barnes & Benjamin, 1924)
Yukon, .... See [About maps] Scotogramma subalbida Barnes & Benjamin, 1924; 123, TL: Alaska, Yukon Territory, Whitehorse
Discestra subalbida ; [POOLE]; [NACL, #10230]
Hadula obesula (Smith, 1904)
Alberta, .... See [About maps] Mamestra obesula Smith, 1904; 151, TL: Alberta, Calgary
Discestra obesula, Butterflies and Moths of North America [BAMONA]
 Discestra obesula ; [NACL, #10231]; [POOLE]
H. o. ortruda (Smith, 1910)
Colorado, ... Mamestra ortruda Smith, 1910; 266, TL: Colorado, Glenwood Springs
= Discestra obesula ; [POOLE]
Discestra obesula ortruda ; [NACL, #10231a]
Hadula crotchi (Grote, 1880)
Oregon, Colorado, .... See [About maps] Mamestra crotchii Grote, 1880; 29, TL: Oregon; Colorado
Discestra crotchii, Butterflies and Moths of North America [BAMONA]
Discestra crotchi, Noctuid Search (record) [NOCTS]
 Discestra crotchi ; [NACL, #10233]
Discestra crotchii ; [POOLE]
H. c. fusculenta (Smith, 1891)
California, ... Mamestra crotchii var. fusculenta Smith, 1891; 213, TL: California
= Discestra crotchii ; [POOLE]
Discestra crotchi fusculenta ; [NACL, #10233a]
Hadula oaklandiae (McDunnough, 1937)
California, .... See [About maps] Scotogramma oaklandiae McDunnough, 1937; 154, f. 2, TL: California, Oakland
Discestra oaklandiae ; [NACL, #10234]; [POOLE]
Hadula chunka (Smith, 1910)
Manitoba, .... See [About maps] Mamestra chunka Smith, 1910; 265, TL: Manitoba, Aweme
Discestra chunka, Butterflies and Moths of North America [BAMONA]
 Discestra chunka ; [NACL, #10235]; [POOLE]
Hadula projecta (McDunnough, 1938)
California, .... See [About maps] Trudestra projecta McDunnough, 1938; 55, f. 1, TL: California, San Bernardino Co., Yermo
Discestra projecta, Butterflies and Moths of North America [BAMONA]
Discestra projecta, Noctuid Search (record) [NOCTS]
 Discestra projecta ; [NACL, #10236]; [POOLE]

 
Hadula agrotiformis (Hampson, 1896)
Ceylon. See [About maps] Hadena agrotiformis Hampson, 1896; 508, TL: Ceylon
Discestra agrotiformis ; [POOLE]
Hadula arenaria (Hampson, 1905)
Pakistan. See [About maps] Discestra arenaria Hampson, 1905; 16, pl. 68, f. 25, TL: Sin, Karachi
Discestra arenaria ; [POOLE]

Larva on Atriplex Wiltshire, 1940, [POOLE]

Hadula basilewskyi (Berio, 1972)
Dicestra basilewskyi Berio, 1972; 290, f. 1, TL: St. Helena Island
Discestra basilewskyi ; [POOLE]
Hadula bergeri (Berio, 1972)
Discerta bergeri Berio, 1972; 291, f. 2, TL: St. Helena Island, Basse Fisher's Valley
Discestra bergeri ; [POOLE]
Hadula quercii (Berio, 1941)
Somalia, .... See [About maps] Scotogramma quercii Berio, 1941; 122, TL: Somalia, Genale
Discestra quercii ; [POOLE]

Unknown or unplaced taxa
Hadena nana var. microdon Guenée, 1852; 96 (repl. Phalaena (Noctua) marmorosa Borkhausen, 1792), TL: "Laponie"
Phalaena (Noctua) marmorosa Borkhausen, 1792; (preocc. Phalaena (Noctua) marmorosa Esper, 1788), TL: "Laponie"
= ; [POOLE]

5.2.2012 (76)

References:
[ALBERTA]
University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection
[About maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[BAMONA]
Paul Opler, Harry Pavulaan, Ray Stanford, Michael Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[ESP]
Heppner, 1981; The dates of E. J. C. Esper's Die Schmetterlinge in Abblidungen... 1776-[1830]; Archives of Natural History (1981) 10 (2): 251-254
[FE]
Fauna Europaea; Fauna Europaea
[Ian Kimber]
http://www.ukmoths.force9.co.uk/
[Karl Hofsäß]
Karl Hofsaess; e-mail: Karl Hofsäß
[Lynn Scott]
Moths of the Ottawa Area; Lynn Scott's Lepidoptera Images
[MBE]
Moths and Butteflies of Europe; Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[NACL]
Hodges, Ronald W. (ed.), 1983; Check List of the Lepidoptera of America North of Mexico
[NE12]
Fibiger, Ronkay, Yela & Zilli, 2010; Rivulinae - Euteliinae, and Micronoctuidae and Supplement to Volume 1-11; Volume 12
[NE4]
Hacker, Ronkay & Hreblay, 2002; Hadeninae I - Noctuidae Europaeae
[NE9]
Fibiger & Hacker, 2007; Amphipyrinae, Condicinae, Eriopinae, Xyleninae (Part) - Noctuidae Europaeae; Volume 9, 410pp
[NOCTS]
Noctuid Search; An Interactive Key to Identify the Noctuidae of North America; e-mail: teichlin@cdfa.ca.gov Thomas D. Eichlin; e-mail: skinnee@cdfa.ca.gov Scott A. Kinnee; Dennis E. Mayhew; Ronald E. Somerby; Charles S. Papp
[POOLE]
Poole, R. W, 1989; Noctuidae; Lepidopterorum catalogus (New Series, Fasc. 118)
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)

Some related literature:
Bang-Haas, 1912
Neue oder wenig bekannte palaearktische Makrolepidopteren. (IV-VI) Dt. ent. Z. Iris 26 (2): 103-110 (1912), (3): 139-162 (1912), (4): 229-230, pl. 6 (1913)
Beck, 1991
Taxonomische Änderungen bei den Noctuinae, Cuculliinae und Plusiinae Atalanta 22 (2/4): 175-232
Boisduval, 1829
Europaeorum Lepidopterorum Index Methodicus Eur. Lepid. Index Meth.: 103pp
Esper, 1787; Esper, 1790
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil IV. Die Eulenphalenen Die Schmett., Th. IV, Bd. 1 (29): 1-16, pl. 80-85 (Noct. 1-6) (1786), (30): 17-32, pl. 86-91 (Noct. 7-12) (1786), (31): 33-48, pl. 92-97 (Noct. 13-18) (1787), (32): 49-64, pl. 98-103 (Noct. 19-24) (1787), (33): 65-80, pl. 104-109 (Noct. 25-30) (1787), (34): 81-96, pl. 110-115 (Noct. 31-36) (1787), (35): 97-112, pl. 116-121 (Noct. 37-42) (1787), (36-37): 113-144, pl. 122-133 (Noct. 43-54) (1788), (38): 145-160, pl. 134-139 (Noct. 55-60) (1788), (39): 161-176, pl. 140-145 (Noct. 61-66) (1789), (40): 177-192, pl. 146-151 (Noct. 67-72) (1790), (41): 193-208, pl. 152-155, 99B, 105B (Noct. 73-76) (1790), (42-44): 209-320, pl. 156-167, 117A (Noct. 77-88) (1791), (45-46): 321-352, pl. 168-177, 125B, 125C (Noct. 89-98) (1791), (47): 353-372, pl. 178-183 (Noct. 99-104) (1794)
Esper, 1804
Die Schmetterlinge in Abbildungen nach der Natur mit Beschreibungen. Theil IV. Die Eulenphalenen Die Schmett., Th. IV, Bd. 2 (Abs. 1) (48): 373-420, pl. 184-186 (Noct. 105-107) (1796), (Abs. 1) (49-50): 421-464, pl. 187-191 (Noct. 108-112) (1798), (Abs. 1) (51): 465-496, pl. [192],[193] (Noct. 113-114) ([1799-1803]), (Abs. 1) (52): 497-536, pl. 194-195 (Noct. 115-116) (1804), (Abs. 1) (53): 537-632, pl. 196 (Noct. 117) (1804), (Abs. 1) (54): 633-698, pl. 197-198 (Noct. 118-119) (1805), (Abs. 2) (49-50): 1-44 (1798), (Abs. 2) (51): 45-54 ([1799-1803]), (Abs. 2) (52): 55-62 (1804), (Abs. 2) (53): 63-85 (1805)
Eversmann, 1837
Kurze notizen ueber einige Schmetterlinge Russlands, als Beiträge zu Treitschke's Supplement zu betrachten. (?2) Bull. Soc. imp. Nat. Moscou 1837 (6) : 29-66
Matsumura, 1927
A list of moths collected on Mt. Daisetsu, with descriptions of new species Ins. Matsumurana 1 (3) : 109-119
Rothschild, 1913
[Expedition to the Central Western Sahara]. Lepidoptera Novit. Zool. 20 (1) : 109-143
Rothschild, 1915
Lepidoptera of the M'Zab country, south Algeria, collected by Dr Ernest Hartert and Carl Hilgert in 1914 Novit. Zool. 22 (2): 228-243
Smith, 1903
New noctuids for 1903, no. 2, with notes on Mamestra and Scotogramma J. N.Y. Ent. Soc. 11 (1) : 1-24, pl. 1
Staudinger, 1889
Centralasiatische Lepidopteren Stettin Ent. Ztg 50 (1-3) : 16-60
Thunberg, 1791
Dissertio Entomologica sistens Insecta Suecica, ... Diss. Ent. sistens Insecta Suecica (1): 1-24 (1 plate) (1784), (2): [25-31] ??], 32-46 (9 unnumbered) (1791), (3): [47],48-52 (2 unnumbered) (1792), (4): 53-62 (1792)
Treitschke, 1835
Die Schmetterlinge von Europa Schmett. Eur. 10 (2): 1-340

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.