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Dryopteris Adanson, 1763
Alvejuuret Wood Ferns Shield Ferns
Dryopteris, Biota of North America Program (query database) [BONAP]
Dryopteris, Den Virtuella Floran [DVF]
 Dryopteris (Polypodiaceae) ; [TSF, 3]; [FPN, 50]

Foodplants for Blasticotoma filiceti filiceti

fiDryopteris filix-mas (L.) Schott, 1834
Kivikkoalvejuuri Male-fern Träjon Echter Wurmfarn Dryoptéris Fougère mâle
Polypodium filix-mas L., 1753
Aspidium filix-mas (L.); Swartz, 1801
Dryopteris filix-mas ; [TFG, 114]; [TSF, #11]; [FPN, 51]; [RK, 49]
[filix-mas-1a.jpg] 783x1044 (~180Kb) Finland, Nokia, Utola, 20.6.2005, Photo © Harri Arkkio [filix-mas-1b.jpg] 902x640 (~118Kb) Finland, Nokia, Utola, 20.6.2005, Photo © Harri Arkkio

fiDryopteris cristata (L.) Gray, 1848
Korpialvejuuri Gray Crested Shield Fern Narrow Swamp Fern Crested Wood Fern Granbräken Kammfarn Dryoptéris à crétes
Polypodium cristatum L., 1753
Aspidium cristatum (L.); Swartz, 1801
Nephrodium cristatum (L.); Michaux, 1803
Dryopteris cristata, Aquatic and Wetland Vascular Plants of the Northern Great Plains [USGS]
Dryopteris cristata, Bilder ur Nordens Flora (in Swedish) [RUNEBERG]
 Dryopteris cristata ; [TFG, 106]; [TSF, #12]; [FPN, 51]; [RK, 49]
Dryopteris clintoniana (Eaton) Dowell, 1906
Broad Swamp Fern Clinton's Wood Fern
Aspidium cristatum v. clintonianum Eaton, 1867
Dryopteris cristata v. clintoniana (Eaton); Underwood, 1893
Dryopteris clintoniana ; [TFG, 108]
Dryopteris celsa (Palmer) Small, 1958
Log Fern
Dryopteris goldiana ssp. celsa Palmer, 1899
Dryopteris celsa ; [TFG, 110]
Dryopteris goldiana (Hooker) Gray, 1848
Giant Wood Fern Goldie's Wood Fern
Aspidium goldianum Hooker, 1822
Nephrodium goldianum (Hooker); Hooker & Greville, 1829
Dryopteris goldiana ; [TFG, 112]
fiDryopteris carthusiana (Vill.) H.P. Fuchs
Metsäalvejuuri Narrow Buckler-fern Spinulose Shield Fern Skogsbräken
Dryopteris spinulosa (Roth); Kunze
Dryopteris carthusiana; [RK, 50]
Dryopteris carthusiana, Aquatic and Wetland Vascular Plants of the Northern Great Plains [USGS]
[carthusiana-1.jpg] 700x571 (~104Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen 		[carthusiana-1x.jpg] 900x601 (~79Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen 		[carthusiana-1y.jpg] 700x583 (~38Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen

Foodplant for Udea decrepitalis


?D. c. -
Toothed Wood Fern Spinulose Wood Fern
Dryopteris lanceolatocristata (Hoffmann); Alston, 1957
Polypodium lanceolatocristata Hoffmann, 1790
Polypodium spinulosum Mueller, 1767-77
 Aspidium spinulosum (Mueller); Swartz, 1801
Dryopteris spinulosa (Mueller); Watt, 1867
Dryopteris austriaca v. spinulosa (Mueller); Fiori, 1943
Dryopteris spinulosa ; Watt, 1867, [TFG, 120 (invalid?)]
fiDryopteris carthusiana × cristata
Dryopteris × uliginosa (A. Braun ex Döll); Druce
Dryopteris carthusiana × cristata ; [RK, 50]
fiDryopteris expansa (C. Presl.) Fraser-Jenkins & Jermy
Isoalvejuuri Nordbräken
Dryopteris assimilis S. Walker
Dryopteris austriaca auct.
Dryopteris dilatata auct.
Dryopteris expansa ; [RK, 50]
fiDryopteris dilatata (Hoffm.) A. Gray, 1848
Etelänalvejuuri Broad Buckler-fern Lundbräken
Polypodium dilatatum Hoffmann, 1795
Dryopteris austriaca auct.
Dryopteris dilatata ; [TFG, 118]; [RK, 50]
D. d. v. (?)
Mountain Wood Fern Spreading Wood Fern
Aspidium spinulosum v. anadenium Robinson, 1907
Dryopteris spinulosa v. americana (Fischer); Fernald, 1915
Aspidium spinulosa v. americanum Fischer, 1848
 Dryopteris campyloptera (Kunze); Clarkson, 1930
Aspidium campylopterum Kunze, 1848
Dryopteris dilata ; [TFG, 118]
Dryopteris austriaca (Jacq.) Woynar
Stachliger Wurmfarn Dryoptéris d'Austriche Spreading Wood-fern
Dryopteris spinulosum Sw.
Dryopteris austriaca ; [TSF, #14]
Drypoteris austriaca ; [FPN, 51]
fiDryopteris fragrans (L.) Schott, 1834
Tuoksualvejuuri Fragrant Shield Fern Fragrant Wood Fern Doftbräken
Polypodium fragrans L., 1753
Dryopteris fragrans ; [TFG, 104]; [RK, 51]
[fragrans.jpg] 732x500 (~146Kb) Above the village Novyi Akatuy, the junction of the mountain ranges Uryumkanskii and Kukul'bei, Aleksandrovskozavodskoi District, E Chita Region, E Transbaikalia, Siberia, Russia. 23rd July 1997, Photo © Oleg Kosterin

D. f. v. remotiuscula (Komarov) Komarov, 1934
Nephrodium fragrans v. remotiusculum Komarov, 1911
Dryopteris fragrans v. remotiuscula ; [TFG, 105 (note)]
Dryopteris villarsii (Bell.) Woynar
Starrer Wurmfarn Dryoptéris de Villars Dryoptéris rigide
Aspidium rigidum Sw.
Dryopteris villarsii ; [TSF, #13]
Dryopteris marginalis (L.) Gray, 1848
Marginal Wood Fern Evergreen Wood Fern Leather Wood Fern
Polypodium marginalis L., 1753
Aspidium marginale (L.); Swartz, 1801
Nephrodium marginale (L.); Michaux, 1803
Dryopteris marginalis ; [TFG, 116]
Dryopteris intermedia (Willdenow) Gray, 1848
Fancy Fern Evergreen Wood Fern Intermediate Wood Fern
Polypodium intermedium Muhlenberg, 1810
Aspidium intermedium Willdenow, 1810
 Dryopteris spinulosa v. intermedia (Willdenow); Underwood, 1893
Dryopteris spinulosa v. intermedia (Willdenow); Morton, 1950
Dryopteris intermedia ; [TFG, 122]
Dryopteris arguta (Kaulf.) Watt.
Coastal Shield-fern
Dryopteris arguta ; [FPN, 51]

 

Other hybrids

Dryopteris celsa × cristata Wherry
Dismal Swamp Fern
Dryopteris × atroplaustris Small, 1938
Dryopteris celsa × cristata ; [TFG, 124]
Dryopteris celsa × intermedia Wherry
Spaced Wood Fern
Dryopteris × separabilis Small, 1938
Dryopteris celsa × intermedia ; [TFG, 124]
Dryopteris clintoniana × cristata Wherry, 1961
Deceptive Wood Fern
Dryopteris clintoniana × cristata ; [TFG, 126]
Dryopteris clintoniana × goldiana Dowell, 1908
Simulated Log Fern
Dryopteris clintoniana × goldiana ; [TFG, 126]
Dryopteris clintoniana × intermedia Dowell, 1908
Dowell's Wood Fern
Filix dowellii Farwell, 1925
Dryopteris × dowellii (Farwell); Wherry, 1961
Dryopteris clintoniana × intermedia Dowell, 1908; TFG, 126
Dryopteris clintoniana × marginalis Slosson, 1910
Dryopteris clintoniana × marginalis ; [TFG, 126]
Dryopteris clintoniana × spinulosa Benedict, 1900
Benedict's Wood Fern
Filix benedictii Farwell, 1923
Dryopteris × benedictii (Farwell); Wherry, 1961
Dryopteris clintoniana × spinulosa ; [TFG, 126]
Dryopteris cristata × goldiana Benedict, 1909
Dryopteris cristata × goldiana ; [TFG, 128]
Dryopteris cristata × intermedia Dowell, 1908
Glandular Swamp Fern Boott's Wood Fern
Aspidium boottii Tuckerman, 1843
Dryopteris × boottii (Tuckerman); Underwood, 1893
Dryopteris cristata × intermedia ; [TFG, 128]
Dryopteris cristata × marginalis (Davenport, 1894)
Miss Slosson's Wood Fern
Aspidium cristatum × marginale Davenport, 1894
Nephrodium slossonae Hahne, 1904
Dryopteris × slossonae (Hahne); Wherry, 1942
Dryopteris cristata × marginalis ; [TFG, 128]
Dryopteris cristata × spinulosa Christensen, 1905
Braun's Wood Fern
Aspidium spinulosum v. uliginosum Braun, 1843
Dryopteris × uliginosa (Braun); Druce, 1909
Dryopteris cristata × spinulosa ; [TFG, 128]
Dryopteris "filix-mas" × goldiana Rugg, 1950
Dryopteris "filix-mas" × goldiana ; [TFG, 130]
Dryopteris "filix-mas" × marginalis Winslow, 1910
Vermont Wood Fern
Dryopteris "filix-mas" × marginalis ; [TFG, 130]
Dryopteris fragrans × intermedia
Dryopteris fragranas v. remotiuscula × spinulosa v. intermedia Tryon, 1942
Dryopteris fragrans × intermedia ; [TFG, 130]
Dryopteris goldiana × intermedia Dowell, 1908
Dryopteris goldiana × intermedia ; [TFG, 130]
Dryopteris goldiana × marginalis Dowell, 1908
Susquehanna Wood Fern
Dryopteris × leedsii Wherry, 1942
Dryopteris goldiana × marginalis ; [TFG, 130]
Dryopteris goliana × spinulosa Benedict, 1909
Poyser's Wood Fern
Nephrodium cristatum clintonianum f. silvaticum Poyser, 1908
Dryopteris × poyseri Wherry, 1960
Dryopteris goliana × spinulosa ; [TFG, 130]
Dryopteris intermedia × marginalis Benedict, 1909
Dryopteris intermedia × marginalis ; [TFG, 132]
Dryopteris intermedia × spinulosa Wherry, 1960
Triploid Wood Fern
Dryopteris × triploidea Wherry, 1960
Nephrodium spinulosum fructuosum Gilbert, 1901
Dryopteris spinulosa v. fructuosa (Gilbert); Trudell, 1930
intermedia f. fructuosa (Gilbert); , 1938
Dryopteris intermedia × spinulosa ; [TFG, 132]
Dryopteris marginalis × spinulosa Slosson, 1904
Pittsford Wood Fern
Dryopteris × pittsfordensis Slosson, 1904
Dryopteris marginalis × spinulosa ; [TFG, 132]

17.9.2005 (14)

References:
[BONAP]
A Synonymized Checklist of the Vascular Flora of the United States, Puerto Rico, and the Virgin Islands; A data base interface; BONAP U.S. Checklist - 1998; Provided by TAMU-BWG; Texas A & M Bioinformatics Working Group; Based on; Biota of North America Program
[DVF]
Den Virtuella floran; http://linnaeus.nrm.se/flora/welcome.html; Ett projekt vid Naturhistoriska riksmuseet (Stockholm, Sweden).; Even for non-Swedish speaker, the distribution maps on species are nice (map buttons "Norden" and "Norra halvklotet")
[FPN]
Hitchcock & Cronquist, 1998; Flora of the Pacific Northwest (11th ed.); ISBN 0-295-95273-3
[Harri Arkkio]
Finland; For the time being these pictures are only for this site. Other usage not allowed.
[Oleg Kosterin]
Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[RK]
Hämet-Ahti, L., Suominen, J., Ulvinen, T. & Uotila, P. (eds), 1998; Retkeilykasvio (Field Flora of Finland); Ed. 4. 656 pp.; Finnish Museum of Natural History, Botanical Museum. Helsinki
[RUNEBERG]
Project Runeberg: Nordic literature online; Project Runeberg; Bilder ur Nordens Fauna (Carl Axel Magnus Lindman)
[TFG]
Edgar T. Therry, ?1995; The Fern Guide: Northeasten and Midland United States and Adjacent Canada; [Original publication 1961]
[TSF]
Eduard Thommen, 1951; Taschenatlas der Schweizer Flora
[USGS]
Northern Prairie Wildlife Research Center; http://www.npwrc.usgs.gov/index.htm

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.